Y-DNA Haplogroup I and its
Subclades - 2016
The entire work is identified by the Version
Number and date given on the
Main Page.
Directions for citing the document are given at
the bottom of the Main
Page.
Because of continuing research, the structure
of the Y-DNA Haplogroup Tree changes and ISOGG
does its best to keep the tree updated with the latest developments
in the field. The viewer may observe other versions of the tree on the
Web. Email
Ray Banks
if the differences need
clarification or if you find any broken links on this page.
^ Indicates a next-generation sequencing
entry which does not yet meet quality guidelines for minimum number of
reads.
^^ Indicates an entry which does not meet quality guidelines but may be
helpful.
~ Indicates only an approximate location on the tree.
The
criteria for a representative SNP printed
in bold for a subclade is: traditional usage,
testing one
in multiple labs, and/or being found in the area of the chromosome used
in recent research studies.
SNPs
listed below in italics
(colored black or red)
are quality variants from next-generation sequencing reports
consistently showing as representing that subgroup.
Contact People for Haplogroup I:
Overall I Contact: Phil
Goff
I1 and Subclades Contact: Steve
Trangsrud
I2a1 P37.2 and Subclades Contact: Zdenko Markovic
I2a2 P217 and Subclades Contact: Aaron Salles Torres
I2b L415 and I2c L596 Subclades Contact: Bob May
I M170/PF3715 ,
CTS48/PF3569, CTS70/PF3570, CTS88/PF3571, CTS646/PF3629,
CTS674/FI1/PF3630, CTS772/PF3631, CTS1006, CTS1301/PF3635,
CTS1555/PF3696, CTS1800/PF3699, CTS2193/PF3703, CTS2387/PF3705,
CTS2514/PF3706, CTS2536/PF3707, CTS3076/PF3712, CTS3383/PF3716,
CTS3384/PF3717, CTS3517/PF3719/V2740 , CTS3641/PF3722,
CTS4077/PF3725, CTS4088/PF3868, CTS4209/PF3726, CTS4239/PF3727,
CTS4272/PF3728, CTS4273/PF3729, CTS4340/PF3730, CTS4637, CTS4664,
CTS4745/PF3734, CTS4752/PF3735, CTS4848/FI7/PF3736, CTS4982/PF3737,
CTS5016/PF3738, CTS5150/PF3739, CTS5263, CTS5622/PF3743,
CTS5650/PF3744, CTS5764/PF3746, CTS5908/PF3747, CTS5946/PF3748,
CTS6231/PF3750, CTS6265/PF3871, CTS6271/PF3751, CTS6334/PF3752,
CTS6343/PF3753, CTS6344/PF3754, CTS6497/PF3756, CTS6751/PF3757,
CTS7026/PF3758, CTS7329, CTS7469, CTS7502/PF3760, CTS7540/PF3761,
CTS7593/PF3763, CTS7831/PF3766, CTS8064/PF3768, CTS8257,
CTS8300/PF3770, CTS8333/PF3771, CTS8345/PF3772, CTS8420/PF3773,
CTS8545/PF3775, CTS8742/PF3776, CTS8876/PF3778, CTS8963,
CTS9264/PF3782, CTS9269/PF3783, CTS9484/PF3785, CTS9618/PF3786,
CTS9838/PF3788, CTS9860/PF3790, CTS10058, CTS10941/PF3838,
CTS11369/PF3840, CTS11441, CTS11540/PF3842, CTS11779/PF3844,
CTS11979/PF3878, FI2/YSC0000281, FI3/L756/PF3662/YSC0000284,
FI4/PF3668, FGC2411/Y1877, FGC2412/Y1909, FGC2413/Y1869, FGC2414/Y1903,
FGC2415/Y1880, FGC2416/S10441/Y1865, FGC2417, FGC2418/Y1841, FGC2419,
FGC2420/Y1907, FGC2421/Y1931, FGC2424, FGC2443/Y1834, FGC2447, FGC2453,
FGC2456, FGC2458/Y1942, FGC2467, FGC3436, FGC7049, FGC7050 ,
L41/PF3787, L503, L578/PF3653, L751/PF3779/YSC0000291,
L755/PF3659/YSC0000283, L758/PF3663/YSC0000285, L772/PF3731/YSC0000263,
L844.1/PF3572.1/YSC0000275.1, L846/PF3646/YSC0000280,
L847/PF3841/YSC0000298, L1197/PF3718/YSC0000260, M258/PF3721,
M11064/PF3634 , P19_1, P19_2, P19_3, P19_4, P19_5, P38,
P212/PF3580, PF3742/U179, PF3574, PF3575, PF3578, PF3585,
PF3586, PF3588, PF3589, PF3590, PF3592, PF3594, PF3596, PF3599, PF3600,
PF3601, PF3602, PF3603, PF3604, PF3605, PF3611, PF3612, PF3616, PF3617,
PF3618, PF3620, PF3622, PF3625, PF3627.2/V218.2, PF3639, PF3640,
PF3641, PF3642, PF3645, PF3649, PF3654, PF3660, PF3661, PF3665, PF3666,
PF3670, PF3672, PF3675, PF3677, PF3685, PF3687, PF3689, PF3694, PF3695,
PF3742/U179, PF3759/YSC0000267, PF3780, PF3794, PF3795, PF3796, PF3797,
PF3799, PF3800, PF3803, PF3804, PF3806, PF3807, PF3809, PF3811, PF3813,
PF3814, PF3815, PF3817, PF3819, PF3822, PF3828, PF3829, PF3833, PF3836,
PF3837, PF3847/YSC0000300, PF3864 , Y1847,
YSC0000256.1, YSC0000272, Z16984, Z16985, Z16986/ZS2286 ,
Z16987
�
I1 M253 ,
AM00084/CTS2044, CTS10/Z2676,
CTS22/Z2677, CTS40/Z2678, CTS136/Z2679, CTS209^/V1092^/Z2680^,
CTS565/Z2700, CTS571/Z2701, CTS641/Z2702, CTS733/Z2703, CTS765^/Z2704^,
CTS883/Z2705, CTS1220/Z2706, CTS1266/Z2707, CTS1393/Z2708,
CTS1739/Z2748, CTS1748/Z2749, CTS1755^/Z2750^, CTS1805/Z2752,
CTS2375^/Z2754^, CTS2410/Z2755, CTS2644/Z2757, CTS2732^/Z2758^,
CTS2738^/Z2759^, CTS2970/Z2760, CTS3268/Z2762, CTS3474/V2712/Z2764,
CTS3506/Z2765, CTS3762/Z2766, CTS3843/Z2767, CTS4037^/V2902^/Z2768^,
CTS4130/V2932/Z2769, CTS4189/Z2770, CTS4295/Z2771, CTS4327/Z2772,
CTS4342/Z2773, CTS4344/Z2774, CTS4387, CTS4400/Z2775, CTS4508/Z2776,
CTS4532^/Z2777^, CTS4612^/Z2778^, CTS4656/Z2779, CTS4675^/Z2780^,
CTS4790/Z2781, CTS5167/Z2784, CTS5247/Z2785, CTS5408/Z2786,
CTS5513^/Z2787^, CTS5643^/Z2790^, CTS5705/Z2791, CTS5783/S63,
CTS5891/Z2793, CTS5993/Z2794,
CTS6006/Z2795, CTS6022/Z2796, CTS6076^/Z2797^, CTS6109/Z2798,
CTS6140/Z2799, CTS6221^/Z2802^, CTS6395/Z2803, CTS6547/Z2804,
CTS6629/Z2805, CTS6765/Z2806, CTS6992, CTS7061/Z2807, CTS7121/Z2808,
CTS7267^/Z2809^, CTS7386/Z2810, CTS7477/Z2811, CTS7652/Z2812,
CTS7751^/Z2813^, CTS7949^/Z2814^, CTS8170/Z2816, CTS8224/Z2817,
CTS8394/Z2818, CTS8652/Z2820, CTS8708/Z2821, CTS8716^/Z2822^,
CTS9010/Z2824, CTS9258/Z2825, CTS9288/Z2826, CTS9368^/Z2827^,
CTS9405/Z2828, CTS9486/Z2829, CTS9487^/Z2830^, CTS9518/Z2831,
CTS9648^/Z2832^, CTS9749/Z2833, CTS9900^, CTS10062/Z2834,
CTS10140/Z2837, CTS10338, CTS10613/Z2840, CTS11036/Z2863, CTS11042/S66,
CTS11091/Z2865,
CTS11126^/Z2866^, CTS11166^/Z2867^, CTS11214/Z2868, CTS11503^/Z2869^,
CTS11526/Z2870, CTS11534/Z2871, CTS11552/Z2872, CTS11751/Z2874,
CTS11775/Z2875, CTS11783/Z2876, CTS11838/Z2879, CTS11950^/Z1880^,
CTS12008/Z2881, F3033.2/M1443.2/V4056.2/Z2838.2, FGC2422/Z2715,
FGC2425/Z2714, FGC2427^/Z2713^, FGC2430/Z2699, FGC2431/Z2710,
FGC2432/V1339/Z2712, FGC2435/Z2698, FGC2436/Y1879, FGC2439/Z2711,
FGC2440/V1427/Y3658, FGC2471/Y1882, FGC2483^/Y1933^,
L64, L75, L80, L81, L118, L121/S62, L123, L124/S64, L125/S65, L157.1,
L186, L187, L495/Z2788, L509/Z2864,
L575/V4059/Z2839, L740^/V3959^/YSC0000296^/Z2836^,
L750/YSC0000292/Z2725, L764/YSC0000271/Z2849, L840,
L841/YSC0000257/Z2751, L842/YSC0000264/Z2789, L843/YSC0000273/Z2878, L845/S110/YSC0000278, L848/YSC0000299/Z2877,
M307.2/P203.2, M450/S109, P30, P40, Page123/Z2763 , S107,
S108, S111/Y1868, S22865/Z2845, V1423/Z2717,
V1771/Z2724, V2264^/Z2728^, Y1886^, YSC0000259/Z2761, YSC0000301/Z2882,
Z2718, Z2719, Z2721, Z2722, Z2723, Z2726, Z2727, Z2729, Z2730^, Z2731,
Z2733, Z2736, Z2737, Z2738, Z2740, Z2741^, Z2742^, Z2743, Z2744, Z2747,
Z2823, Z2841, Z2842, Z2843, Z2844, Z2846, Z2847, Z2848, Z2850, Z2851^,
Z2852^, Z2853^, Z2854^, Z2855^, Z2856, Z2857^, Z2861, Z2862, Z2885,
Z2887
� I1~
L234, L410, L1274, L1275
� �
I1a DF29/S438
� � �
I1a1 CTS6364/Z2336
� � � �
I1a1a M227
� � � � �
I1a1a1 M72
� � � �
I1a1b L22/S142
� � � � I1a1b~ S336/Z75
� � � � �
I1a1b1 P109
� � � � �
I1a1b2 L205.1/L939.1/S239.1
� � � � �
I1a1b3 Z74
� � � � �
�
I1a1b3a L287/S240
� � � � �
� �
I1a1b3a1 L258/S335
� � � � �
� � �
I1a1b3a1a L296
� � � � �
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I1a1b3b L813/S436/Z719
� � � � �
I1a1b4 L300/S241
� � � � �
I1a1b5~ S297/Z718
� � � � �
I1a1b6~ L242
� � �
I1a2 S244/Z58
� � � �
I1a2a S246/Z59
� � � � �
I1a2a1 S337/Z60 ,
S439/Z61, Z62
� � � � �
�
I1a2a1a S440/Z140 ,
Z141
� � � � �
� �
I1a2a1a1 S1953/Z2535
� � � � �
� � �
I1a2a1a1a S1954/YSC0000261
� � � � �
� � � �
I1a2a1a1a1 L338/ S197
� � � � �
� � � � I1a2a1a1a1~
A376
� � � � �
� � �
I1a2a1a1b CTS10937/S4698/Z2538
� � � � �
� �
I1a2a1a2 F2642.1/S2169.1
� � � � �
� �
I1a2a1a3 A196
� � � � �
� � �
I1a2a1a3a Y6900
� � � � �
� � � �
I1a2a1a3a1 Y7140
� � � � �
� � � � �
I1a2a1a3a1a Y7477
� � � � �
� � � � �
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I1a2a1a3a1a1 YP1081
� � � � �
� � � �
I1a2a1a3a2 Y6885
� � � � �
�
I1a2a1b S247/Z73
� � � � �
� �
I1a2a1b1 BY266/Y3657
� � � � �
� � �
I1a2a1b1a L1302
� � � � �
� � � I1a2a1b1a~
BY148, BY211, BY212, BY213, BY214, BY215, BY216, BY217, BY218, BY220,
BY222, BY223, BY224, BY225, BY226, BY228, BY230, BY232, BY233, BY234,
BY235, BY236, BY240, BY244, BY245, BY269, BY270, BY271, BY272, BY273,
BY274, BY275, BY276, BY277, BY278
� � � � �
� � � �
I1a2a1b1a1 BY147
� � � � �
� � � �
I1a2a1b1a2 BY126
� � � � �
�
I1a2a1c L573
� � � � �
�
I1a2a1d L1248
� � � � �
� �
I1a2a1d1 L803
� � � � �
� I1a2a1e~
CTS7362/S4713/Z2539
� � � � �
I1a2a2 Z382
� � � � � I1a2a2~
L1450, L1451, L1452, Z2037, Z2038, Z2040, Z2041
� � � �
I1a2b S296/Z138 ,
Z139/S338, PF2805.2/Z2540.2
� � � � �
I1a2b1 Z2541
� � �
I1a3 S243/Z63
� � � I1a3~
BY12, BY13
� � � �
I1a3a BY151
� � � � �
I1a3a1 L849.2
� � � � �
�
I1a3a1a S2078
� � � � �
� �
I1a3a1a1 Y2245.2
� � � � �
� � �
I1a3a1a1a L1237
� � � � �
� � � �
I1a3a1a1a1 FGC9550/Y6634
� � � � �
� � �
I1a3a1a1b S10360 ,
S17023, S26266
� � � � �
� � � �
I1a3a1a1b1 S15301
� � � � �
� � � � �
I1a3a1a1b1a Y6228
� � � � �
� � � � �
�
I1a3a1a1b1a1 Y6229
� � � � �
� � � � �
I1a3a1a1b1b Y3969
� � � � �
� � �
I1a3a1a1c CTS3848 ,
FGC14479
� � � � �
� �
I1a3a1a2 Y6375 ,
Y6383
� � � � �
I1a3a2 BY351 ,
BY352
� � � � �
�
I1a3a2a BY332 ,
BY354/ Y7075
� � � � �
� �
I1a3a2a1 Y7060 ,
Y7059
� � � � �
� � �
I1a3a2a1a BY11 ,
S11542.1/V2110.1
� � � � �
� �
I1a3a2a2 Y13946 ,
PH1538/Y14110
� � � � �
�
I1a3a2b CTS10345 ,
CTS2910/S4736
� � � � �
� �
I1a3a2b1 Y10994
� � � � �
�
I1a3a2c CTS4279
� � � � �
I1a3a3 BY62
� �
I1b S249/Z131
� � �
I1b1 CTS6397
� �
I1c Y18119/Z17925 ,
S2304/Z17937, Z17943
�
I2 M438/P215/PF3853/S31 ,
L68/PF3781/S329, PF3664/Z2638
� �
I2a L460/PF3647/S238
� � �
I2a1 P37.2/ PF4004
� � � �
I2a1a CTS595
� � � � �
I2a1a1 M26/PF4056 ,
L158/PF4073/S433,
L159.1/S169.1
� � � � �
I2a1a1~ L673
� � � � �
�
I2a1a1a L672/S327
� � � � �
� �
I2a1a1a1 L160/PF4013/S184
� � � � �
� � I2a1a1a1~
Z109, Z112, Z124
� � � � �
� � �
I2a1a1a1a PF4088/S5312
� � � � �
� � � �
I2a1a1a1a1 CTS11338/PF4194
� � � � �
� � � � �
I2a1a1a1a1a PF4189
� � � � �
� � � � �
I2a1a1a1a1b Z118
� � � � �
� � � � �
� I2a1a1a1a1b1~
S294/Z106
� � � � �
� � �
I2a1a1a1b F1295
� � � � �
� � � I2a1a1a1c~
Z119, Z125
� � � � �
I2a1a2 S21825
� � � � �
�
I2a1a2a L1286
� � � � �
� �
I2a1a2a1 L1287
� � � � �
� � �
I2a1a2a1a L233/S183
� � � � �
� � � �
I2a1a2a1a1 A417
� � � � �
� �
I2a1a2a2 L880
� � � � �
�
I2a1a2b L1294
� � � �
I2a1b M423 ,
CTS176/S2621, CTS1293/S2632, CTS1802/S2638, CTS5375/S2679,
CTS5985/S2687, CTS7218/S2702, CTS8239/S2715, CTS8486/S2722,
CTS11030/S2768, L178/S328
� � � � I2a1b~ L1224
� � � � �
I2a1b1 L161.1/S185
� � � � �
�
I2a1b1a
AMM079.1 /L1498.1/Y4008.1
� � � � �
� �
I2a1b1a1 S2703
� � � � �
�
I2a1b1b PF4135
� � � � �
� I2a1b1b~
A813, A814
� � � � �
I2a1b2 L621/S392
� � � � �
�
I2a1b2a CTS10936
� � � � �
� �
I2a1b2a1 CTS5966 ,
CTS10228, L147.2
� � � � �
� � I2a1b2a1~
A493
� � � � �
� � �
I2a1b2a1a S17250/YP204
� � � � �
� � � �
I2a1b2a1a1 Z16971
� � � � �
� � � � I2a1b2a1a1~
BY128
� � � � �
� � � � � I2a1b2a1a1a~ A815
� � � � �
� � � �
I2a1b2a1a2 Y4882
� � � � �
� � � � � I2a1b2a1a2a~
A811
� � � � �
� � � �
I2a1b2a1a3 A356/Z16983
� � � � �
� � � � I2a1b2a1a4~
M359.2/P41.2
� � � � �
� � �
I2a1b2a1b Y4460
� � � � �
� � � �
I2a1b2a1b1 Y3118
� � � � �
� � �
I2a1b2a1c Z17855
� � �
I2a2 M436/P214/PF3856/S33 ,
L35/PF3862/S150,
L37/PF6900/S153, L181, L368, P216/PF3855/S30, P217/PF3854/S23, P218/S32
� � � �
I2a2a M223 ,
L34/PF3857/S151,
L36/S152, L59, L622, P219/PF3859/S24, P220/S119, P221/PF3858/S120,
P222/PF3861/U250/S118,
P223/PF3860/S117, Z77
� � � � I2a2a~
L387.2, L700, L705.1, L706.1
� � � � �
I2a2a1 CTS616 ,
CTS9183
� � � � �
�
I2a2a1a FGC15073 /Y3721
� � � � �
� �
I2a2a1a1 M284
� � � � �
� � �
I2a2a1a1a L1195
� � � � �
� � � �
I2a2a1a1a1 L126/S165 ,
L137.1/S166.1, L369
� � � � �
� � � � �
I2a2a1a1a1a FGC20048/S7753/Y4171
� � � � �
� � � � �
�
I2a2a1a1a1a1 Y8701
� � � � �
� � � � �
� �
I2a2a1a1a1a1a Y4751
� � � � �
� � � � �
� � �
I2a2a1a1a1a1a1 Y10655
� � � � �
� � � � �
� � �
I2a2a1a1a1a1a2 Y4752
� � � � �
� � � � �
� � � �
I2a2a1a1a1a1a2a Y4750
� � � � �
� � � � �
� � � � �
I2a2a1a1a1a1a2a1 A824/Y11197
� � � � �
� � � � �
� � � � �
I2a2a1a1a1a1a2a2 Y5673
� � � � �
� � � � �
� � � � �
�
I2a2a1a1a1a1a2a2a Y11246 ,
S15280.2
� � � � �
� � � � �
� � � � �
I2a2a1a1a1a1a2a3 Y8599
� � � � �
� � � � �
� �
I2a2a1a1a1a1b Y7190
� � � � �
� � � �
I2a2a1a1a2 L1193
� � � � �
� � � � �
I2a2a1a1a2a CTS4922 ,
Y3684, Y3713
� � � � �
� �
I2a2a1a2 Z2057 ,
Z2083, Z2093
� � � � �
� � �
I2a2a1a2a L1229
� � � � �
� � � �
I2a2a1a2a1 Y3681 ,
Z2069
� � � � �
� � � � �
I2a2a1a2a1a Z2058
� � � � �
� � � � �
�
I2a2a1a2a1a1 Z2054
� � � � �
� � � � �
� I2a2a1a2a1a1~
BY18
� � � � �
� � � � �
� �
I2a2a1a2a1a1a BY524/Y9443
� � � � �
� � � � �
� � �
I2a2a1a2a1a1a1 L812/S391
� � � � �
� � � � �
� � � �
I2a2a1a2a1a1a1a Y5308 ,
Y5318
� � � � �
� � � � �
� � � �
I2a2a1a2a1a1a1b Y13008
� � � � �
� � � � �
� � �
I2a2a1a2a1a1a2 Y10648
� � � � �
� � � � �
� �
I2a2a1a2a1a1b Y7244
� � � � �
� � � � �
� � �
I2a2a1a2a1a1b1 P53.3
� � � � �
� � � � �
� � �
I2a2a1a2a1a1b2 Y7243
� � � � �
� � � � �
� �
I2a2a1a2a1a1c FGC15106/Y4334 ,
Y4332
� � � � �
� � � � �
� � �
I2a2a1a2a1a1c1 Y4760
� � � � �
� � � � �
� �
I2a2a1a2a1a1d BY138
� � � � �
� � � � �
� � �
I2a2a1a2a1a1d1 Y13321
� � � � �
� � � �
I2a2a1a2a2 S18331 ,
SK1247/Y4915
� � � � �
� � � � �
I2a2a1a2a2a L1230
� � � � �
� � �
I2a2a1a2b Y7240
� � � � �
�
I2a2a1b CTS10057 ,
CTS10100
� � � � �
� �
I2a2a1b1 L701 ,
L702
� � � � �
� � �
I2a2a1b1a P78
� � � � �
� � � �
I2a2a1b1a1 S25733
� � � � �
� � � � �
I2a2a1b1a1a A427
� � � � �
� � � � �
�
I2a2a1b1a1a1 S23612 ,
Y4884
� � � � �
� � � � �
� �
I2a2a1b1a1a1a Y5360 ,
Y5359
� � � � �
� � � � �
� � �
I2a2a1b1a1a1a1 Y5369
� � � � �
� � � � �
� �
I2a2a1b1a1a1b S10702 ,
Y6396
� � � � �
� � � � �
� � �
I2a2a1b1a1a1b1 Y11261
� � � � �
� � � �
I2a2a1b1a2 Y7219
� � � � �
� � � � �
I2a2a1b1a2a Y7214
� � � � �
� � � � �
I2a2a1b1a2b Y8945
� � � � �
� � �
I2a2a1b1b L699 ,
L703
� � � � �
� � � �
I2a2a1b1b1 L704
� � � � �
� � � � �
I2a2a1b1b1a Y7641
� � � � �
� � � �
I2a2a1b1b2 S12195 ,
S21579, Y5669
� � � � �
� � � � �
I2a2a1b1b2a Y6973 ,
BY159/Y6996
� � � � �
� �
I2a2a1b2 Z161
� � � � �
� � �
I2a2a1b2a L801/S390
� � � � �
� � � �
I2a2a1b2a1 CTS1977
� � � � �
� � � � �
I2a2a1b2a1a Y5282
� � � � �
� � � � �
�
I2a2a1b2a1a1 S8522
� � � � �
� � � � �
� �
I2a2a1b2a1a1a P95
� � � � �
� � � � �
�
I2a2a1b2a1a2 CTS1858
� � � � �
� � � � �
� �
I2a2a1b2a1a2a CTS10148
� � � � �
� � � � �
I2a2a1b2a1b Y7152 ,
S16196
� � � � �
� � � � �
I2a2a1b2a1c BY526/Y8935
� � � � �
� � � �
I2a2a1b2a2 CTS6433
� � � � �
� � � � �
I2a2a1b2a2a S2364
� � � � �
� � � � �
�
I2a2a1b2a2a1 S2361 ,
Z167
� � � � �
� � � � �
� �
I2a2a1b2a2a1a Z78
� � � � �
� � � � �
� � �
I2a2a1b2a2a1a1 CTS8584
� � � � �
� � � � �
� � � �
I2a2a1b2a2a1a1a Z185
� � � � �
� � � � �
� � � � �
I2a2a1b2a2a1a1a1 Z180
� � � � �
� � � � �
� � � � �
�
I2a2a1b2a2a1a1a1a L1198
� � � � �
� � � � �
� � � � �
� �
I2a2a1b2a2a1a1a1a1 S20905
� � � � �
� � � � �
� � � � �
� � �
I2a2a1b2a2a1a1a1a1a Z190
� � � � �
� � � � �
� � � � �
� � � �
I2a2a1b2a2a1a1a1a1a1 S434/Z79
� � � � �
� � � � �
� � � � �
� � � � �
I2a2a1b2a2a1a1a1a1a1a Y5729 ,
Y5730
� � � � �
� � � � �
� � � � �
� � � � �
I2a2a1b2a2a1a1a1a1a1b PH3769/Y7682
� � � � �
� � � � �
� � � � �
� � � � �
I2a2a1b2a2a1a1a1a1a1c Y8712
� � � � �
� � � � �
� � � � �
� � � �
I2a2a1b2a2a1a1a1a1a2 Y7280
� � � � �
� � � � �
� � � � �
� �
I2a2a1b2a2a1a1a1a2 P195.2
� � � � �
� � � � �
� � � � �
� �
I2a2a1b2a2a1a1a1a3 Y6060
� � � � �
� � � � �
� � � � �
� � �
I2a2a1b2a2a1a1a1a3a Y5748
� � � � �
� � � � �
� � � � �
� � � �
I2a2a1b2a2a1a1a1a3a1 Y7272
� � � � �
� � � � �
� � � � �
� � �
I2a2a1b2a2a1a1a1a3b Y7273
� � � � �
� � � � �
� �
I2a2a1b2a2a1b Y9161
� � � � �
� � � � �
�
I2a2a1b2a2a2 ZS20/Y4925 ,
S8112
� � � � �
� � � � �
� �
I2a2a1b2a2a2a CTS661/L1272
� � � � �
� � � � �
� � �
I2a2a1b2a2a2a1 Y5717
� � � � �
� � � � �
� �
I2a2a1b2a2a2b Y4926
� � � � �
� � � � �
� � �
I2a2a1b2a2a2b1 S8104 ,
Y6883
� � � � �
� � � � �
�
I2a2a1b2a2a3 Y4955
� � � � �
� � � � �
� �
I2a2a1b2a2a3a Y5695
� � � � �
� � � � �
� � �
I2a2a1b2a2a3a1 Y7263
� � � � �
� � � � �
� � � �
I2a2a1b2a2a3a1a Y7265
� � � � �
� � � � �
� � �
I2a2a1b2a2a3a2 Y5692
� � � � �
� � � � �
� �
I2a2a1b2a2a3b Y10659
� � � � �
� � � � �
I2a2a1b2a2b S25383 ,
L1425
� � � � �
� � � � �
�
I2a2a1b2a2b1 CTS5332.2/PF7472.2
� � � � �
� � � � �
�
I2a2a1b2a2b2 Y4769
� � � � �
� � � � �
I2a2a1b2a2c Y7426
� � � � �
� � �
I2a2a1b2a3 L1290 ,
L1317
� � � � �
� � �
I2a2a1b2a4 Y7202 ,
Y7844
� � � � �
� � �
I2a2a1b2b L623 ,
L147.3
� � � � �
� � �
I2a2a1b2c CTS11871/S3673
� � � � �
� � � �
I2a2a1b2c1 Y10671
� � � � �
I2a2a2 S9403/SK1254 ,
Y6098
� � � � �
�
I2a2a2a L1228
� � � � �
�
I2a2a2b Y6099
� � � �
I2a2b L38/S154 ,
L39/S155, L40/S156,
L65.1/S159.1, L272.3
� � � � �
I2a2b1 L533/S295
� � � � �
I2a2b2 S2606
� � � � �
�
I2a2b2a S24121
� � � � �
� �
I2a2b2a1 Y16415
� � � � �
� � �
I2a2b2a1a Y16417
� � � � �
� � �
I2a2b2a1b Y17117
� � � � �
�
I2a2b2b FGC29612^^/Y13070^^
� � � � �
� �
I2a2b2b1 S2488 ,
Y13074
� � � � �
� �
I2a2b2b2 PH1237
� � � � �
� �
I2a2b2b3 Y18919
� � � � �
� � �
I2a2b2b3a Y18921
� � � � �
� � � �
I2a2b2b3a1 Y19084
� � � � �
� � � � �
I2a2b2b3a1a Y19211
� � � � �
� � � � �
I2a2b2b3a1b Y20202
� � I2b~ L415/S435, L416, L417/S332
� �
I2c L596/PF6907/S292 ,
L597/S333,
FGC18085/Y5340, FGC18140/Y5336, M3934/S6610/PF3934, PF3889, PF3893,
PF3895, PF3896, PF3897, PF3898/S6659, PF3903/S3330, PF3911/S6728,
PF3922, PF3929/S6694, PF3931, S6601, S6631/SK1266, S11759, S12355,
Y5332, Y5338
� � I2c~
BY19, BY20, BY21, BY22/FGC18100, BY24, BY25, BY26, BY27, BY29,
PF3881/S6673
� � �
I2c1 PF3892 ,
PF3904, PF3918, PF3924, PF3933
� � � �
I2c1a BY433
� � � � �
I2c1a1 L1251.1 ,
BY2776, BY2777, S9234, S10141, S11571, S16199, S16446, S17221,
S17911, S18042, S18299, S21750, S22170, S22470, S23078, S26491
� � � � �
�
I2c1a1a FGC18564 ,
S14736
� � � � �
� �
I2c1a1a1 BY2789 ,
BY2790, BY2791, BY2792
� � � � �
I2c1a2 S6687 ,
S6741
� � � � �
�
I2c1a2a CTS4092
� � � � �
� �
I2c1a2a1 CTS7767.1
� � � � �
� � �
I2c1a2a1a BY23/S6625
� � � � �
� � � �
I2c1a2a1a1 PF6328
� � � � �
� � � � �
I2c1a2a1a1a F2044
� � �
I2c2 BY133.1^/Y5127.1^ ,
PH3539, Y16659, Y16660, Y16668, Y16672, Y16673, Y16675, Y16677, Y16678,
Y16680, Y16682, Z26376, Z26380, Z26383, Z26384^, Z26386, Z26388,
Z26389, Z26390, Z26396^, Z26399, Z26401, Z26403^, Z26404
� � � I2c2~ Y16427
� � � �
I2c2a Z26381
� � � � �
I2c2a1 Y16666
� � � � �
�
I2c2a1a FGC6702.2/Y14376.2
� � � � �
� I2c2a1b~
Y16423
Notes
Identical SNPs that were discovered separately are listed
in alphabetical order, not necessarily in the order of discovery, and
separated by "/". Examples: M307.1/P203.1, M450/S109, M438/P215,
P41.2/M359.2, M436/P214/S33, P222/U250, L126/S165, L137/S166, L38/S154,
L39/S155, L40/S156, and L65/S159.
Page76: published accounts place this downstream from M170,
which appears to be incorrect. A haplogroup J man has tested Page76- as
expected, but three I1 males were Page76- as well. No known I male has
been tested for Page76. Therefore Page76 is possibly in I2, but
definitely not at the I node.
Page123: The precise location of Page123 is uncertain. It
is derived in four men with Haplogroup I1 SNPS and ancestral for one
man in Haplogroup J. It has not been tested in Haplogroup I2.
L69/S163 - Removed from I in 2011 and IJK in 2012,
frequently mutating.
Mutations previously on tree but found so far
only in one man or in closely related men :
S79 : One
confirmed results downstream from P30; considered a private snp, was
previously listed as I1e. Listed 26 Sep 2008.
L592 : Out of
15 men tested, only three, all with the Straub surname, were derived.
This snp is downstream from Z140/Z141. Listed February 15, 2012.
L211 :
Downstream from L64 et al. As of 16 April 2012, 219 men have tested
from L211, of which only two of the same surname have been confirmed as
L211+
M21 :One
confirmed tester in Underhill et al (197), downstream for L64 et al. As
of 17 April 2012, 969 men in the FTDNA I1 Project have tested and all
are ancestral.
M507/P259 :
One confirmed result in Karafet et al (2008) downstream from L64 35 al.
As of 17 April 2012, 679 men in the FTDNA I1 Project have tested and
all are ancestral.
M379 :
Downstream of M223, only one instance of M379+ is known, a Pakistani
male in Polarity and Temporality of High-Resolution
Y-ChromosomeDistributions in India Identify Both Indigenous and
Exogenous Expansions and Reveal Minor Genetic Influence of Central
Asian Pastoralists by Sengupta et al. The Pakistani sample was derived
for M223. In the FTDNA I2b1/M223 Project, 237 men have tested for M379
and all were ancestral. Listed 28 June 2012.
M161/PF4447
is downstream of L160. Listed 5 August 2012.
PF6479 is
approximately PF3827. Listed 18 December 2014, but PF3827 does not
exist on tree.
Y-DNA haplogroup I is a European
haplogroup, representing nearly one-fifth of the
population. It is almost non-existent outside of Europe, suggesting
that it arose in Europe.
Estimates of the age of haplogroup I suggest that it arose prior to the
last Glacial Maximum.
The two main subgroups of haplogroup I likely divided
approximately 28,000 years ago:
I1-M253 et al has highest frequency in Scandinavia,
Iceland, and northwest Europe. In Britain, haplogroup I1-M253 et al is
often used as a marker for "invaders," Viking or Anglo-Saxon. The
I1b-M227 subclade is concentrated in eastern Europe and the Balkans and
appears to have arisen in the last one thousand to five thousand years.
It has been reported in Germany, Czech Republic, Poland, Estonia,
Ukraine, Switzerland, Slovenia, Bosnia, Macedonia, Croatia, and Lebanon.
I2-M438 et al includes I2* which shows some membership from
Armenia, Georgia and Turkey; I2a1-P37.2, which is the most common form
in the Balkans and Sardinia. I2a1a-M26 is especially prevalent in
Sardinia. I2a2-M436 et al reaches its highest frequency along the
northwest coast of continental Europe. I2a2a-M223 et al occurs in
Britain and northwest continental Europe. I2a2a1-M284 occurs almost
exclusively in Britain, so it apparently originated there and has
probably been present for thousands of years.
References:
Athey et al,
Resolving the
Placement of Haplogroup I-M223 in the Y-chromosome Phylogenetic Tree.
(pdf)
Journal of Genetic Genealogy, 1:54-55, 2005.
Battaglia et al,
Y-chromosomal
Evidence
of the Cultural Diffusion of Agriculture in Southeast Europe
European Journal of Human Genetics, 249. 2008.
Behar et al,
Genome-Wide
Structure of the Jewish People.
Nature, 446:238-42, 2010.
Bosch et al,
Paternal
and Maternal Lineages in the Balkans Show a Homogeneous Landscape over
Linguistis Barriers
except for the Isolated Aromuns.
Annals of Human Genetics, 70:459-87, (2006).
Cinnioglu et al,
Excavating Y-chromosome Haplotype Strata in Anatolia. (pdf)
Human Genetics. 114:127-148, 2004.
El Sibai et al,
Geographical
Structure of the Y-Chromosomal Genetic Landscape of the Levant: A
Coastal Inland
Contrast , Annals of Human Genetics, 73:568-81, 2009.
(abstract)
Francalacci et al,
Low-Pass
DNA Sequencing of 1200 Sardinians Reconstructs European Y-Chromosome
Phylogeny.
Science: Vol. 341 no. 6145, pp. 565-569, DOI: 10.1126/science.1237947,
2 August 2013.
Ghiani , et al,
Population
data for Y-chromosome haplotypes defined by AmpFlSTR YFiler PCR
amplification kit in
North Sardinia (Italy) , Collegium Antropologicum, 33 (2):
643-51, 2009.
Karafet et al,
New
Binary Polymorphisms Reshape and Increase Resolution of the Human
Y-Chromosomal Haplogroup
Tree. Abstract. Genome Research, published online April 2,
2008.
Supplementary
Material.
Rootsi et al,
Phylogeography
of Y-Chromosome Haplogroup I Reveals Distinct Domains of Prehistoric
Gene
Flow In Europe. American Journal of Human Genetics,
75:128-137, 2004.
Rozen et al,
Remarkably
Little Variation in Proteins Encoded by the Y Chromosome's Single-Copy
Genes, Implying
Effective Purifying Selection. American Journal of Human
Genetics, 85:923-928, 2009.
Scheinfeldt et al,
Unexpected
NRY Chromosome Variation in Northern Island Melanesia.
Society for Molecular Biology, 2006.
Sims et al,
Sub-Populations Within the Major European and African Derived
Haplogroups R1b3 and
E3a Are Differentiated by Previously Phylogenetically Undefined Y-SNPs.
Human Mutation: Mutation in Brief #940, Online, 2007.
Underhill et al,
New Phylogenetic Relationships for Y-chromosome Haplogroup I:
Reappraising its Phylogeography and Prehistory
in Rethinking the Human Evolution ,
Mellars P, Boyle K, Bar-Yosef O, Stringer C, Eds.
McDonald Institute for Archaeological Research, Cambridge, UK, pp.
33-42, 2007b. (not available online)
Underhill et al,
The Phylogeography of Y Chromosome Binary Haplotypes and the Origins of
Modern Human
Populations. (pdf) Annals of Human Genetics, 65:43-62, 2001.
Underhill et al,
Y Chromosome Sequence Variation and the History of Human Populations.
(pdf) Nature Genetics, 26:
358-361, 2000.
Y Chromosome Consortium (YCC) ,
A Nomenclature System for the Tree of Human Y-Chromosomal Binary
Haplogroups. Genome
Research, 12:339-348, 2002.
Additional Resources:
ISOGG
Wiki - What you need to know about Genetic Genealogy.
Founder
Haplotypes (Modal Haplotypes for Y-Haplogroup I
Varieties) , Ken Nordtvedt.
Deep
Ancestry of I1a DYS19=16
Cluster , Phil Goff.
I1b*
Project Database , Timothy Weakley.
Family Tree DNA Projects:
Haplogroup I Y-DNA Project , Bob Waterous, Bernie Cullen, Paul
Stone.
I1
Y-DNA Haplogroup Project ,
Phil Goff, Aaron J. Hill, Steve Transgrud, Yme Drost, Ken Nordtvedt,
William Hartley and Paul Stone.
I1>Z58+
and I1>Z63+ Project , Aaron Salles Torres,
Paul Stone, William Hartley, Wim Penninx.
I1d
- L22/S142 Project , Yme Drost, Albert Hulse.
I1d2
(L205+) Project , Yme Drost.
The
I1-L1302 Project , Peter Sj�lund, Jakob Norstedt-Moberg
I1-Z140
YDNA Project , William Hartley, Wim Penninx and Aleksandr
Shtrunov.
Y-Haplogroup
I2a Project , Bernie Cullen, Bill Morrow, Ken Nordtvedt,
Zdenko Markovic.
I2b2
L38+ Project , Timothy Weakley, Hans De Beule, Ken Nordtvedt.
I2b
L415 I2c L596 Haplogroup Project , Russ Livingstone, Bob May,
Peter Hrechdakian, Ken Nordtvedt, Hovann Simonian.
Iberian
I1 Project , Aaron Salles Torres.
I-L161
(I2a-Isles) , Robert Gabel.
I-M223
Y-Haplogroup Project , Aaron Salles Torres, Wayne Roberts, Ken
Nordtvedt, George Anthony, Scott Engel Stewart.
The
I1d1 (I-P109) yDNA Haplogroup Project , Rebekah Canada,
Nicolas Taban.
The
I1-L300 and Subclades yDNA Haplogroup Project , Artem Chuprov.
The
I-L1248 and Subclades yDNA Haplogroup Project , Jakob
Norstedt-Moberg.
I1
Eastern Europe , Aleksandr Shtrunov.
Corrections/Additions made since 1 January 2016:
Moved BY211, BY212, BY213, BY214, BY215, BY216, BY217,
BY218, BY220, BY222, BY223, BY224, BY225, BY226, BY228, BY230, BY232,
BY233, BY234, BY235, BY236, BY240, BY244, BY245, BY269, BY270, BY271,
BY272, BY273, BY274, BY275, BY276, BY277, BY278, BY19, BY20, BY21,
BY22/FGC18100, BY24, BY25, BY26, BY27, BY29, BY148, BY18, BY128, BY12,
BY13, A815, A813, A814, A811, A493, A376, M359.2/P41.2, PF3881/S6673,
L1450, L1451, L1452, Z2037, Z2038, Z2040, L96/S196, L1224,
CTS7362/S4713/Z2539, L1274, L1275, Z2041, Z119, Z125, S294/Z106, Z109,
Z112, Z124, L673, L387.2, L700, L705.1/S465.1, L706.1, S336/Z75,
S297/Z718, L234, L410, L242 from SNPs under Investigation to Tree
Investigation; moved PF5479 from SNPs under Investigation to Notes;
removed L96/S196 has nonsensical information on 7 January 2016.
Added BY133.1/Y5127.1, PH3539, Y16659, Y16660, Y16668,
Y16672, Y16673, Y16675, Y16677, Y16678, Y16680, Y16682, Z26376, Z26380,
Z26383, Z26384, Z26386, Z26388, Z26389, Z26390, Z26396, Z26399, Z26401,
Z26403, Z26404 on 13 January 2016.
Added PF3664/Z2638, Z26381, Y16666, Y14976.2/FGC6702.2 to
tree on 14 January 2016.
Added Y16423, Y16427 to tree investigation on 17 January
2016.
Added V2110.1 to tree on 14 February 2016.
Removed CTS3529/PF3720/Z2649, CTS4039/PF3723/Z2650,
CTS8959/Z2659, CTS9380/PF3784, PF3801/Z2663, PF3802/Z2664 from tree due
to erroneous entries; removed BY417 from tree not meeting quality
guidelines on 17 February 2016; moved L415/S435, L416, L417/S332 from
tree to tree investigation.
Removed S465.1 wrong mutation information for haplogroup I
on 20 Mar 2016.
Added Y16415, Y16417, Y17117 to tree on 5 April 2016.
Added FGC29612/Y13070, PH1237, Y18919, Y18921, Y19084,
Y19211, Y20202 to tree on 12 April 2016.
Added AM00084/CTS2044, CTS10/Z2676, CTS22/Z2677,
CTS40/Z2678, CTS136/Z2679, CTS209/V1092/Z2680, CTS565/Z2700,
CTS571/Z2701, CTS641/Z2702, CTS733/Z2703, CTS765/Z2704, CTS883/Z2705,
CTS1220/Z2706, CTS1266/Z2707, CTS1393/Z2708, CTS1739/Z2748,
CTS1748/Z2749, CTS1755/Z2750, CTS1805/Z2752, CTS2375/Z2754,
CTS2410/Z2755, CTS2644/Z2757, CTS2732/Z2758, CTS2738/Z2759,
CTS2970/Z2760, CTS3268/Z2762, CTS3474/V2712/Z2764, CTS3506/Z2765,
CTS3762/Z2766, CTS3843/Z2767, CTS4037/V2902/Z2768, CTS4130/V2932/Z2769,
CTS4189/Z2770, CTS4295/Z2771, CTS4327/Z2772, CTS4342/Z2773,
CTS4344/Z2774, CTS4387, CTS4400/Z2775, CTS4508/Z2776, CTS4532/Z2777,
CTS4612/Z2778 to tree on 25 April 2016.
Added CTS4656/Z2779, CTS4675/Z2780, CTS4790/Z2781,
CTS5167/Z2784, CTS5247/Z2785, CTS5408/Z2786, CTS5513/Z2787,
CTS5643/Z2790, CTS5705/Z2791, CTS5891/Z2793, CTS5993/Z2794,
CTS6006/Z2795, CTS6022/Z2796, CTS6076/Z2797, CTS6109/Z2798,
CTS6140/Z2799, CTS6221/Z2802, CTS6395/Z2803, CTS6547/Z2804,
CTS6629/Z2805, CTS6765/Z2806, CTS6992, CTS7061/Z2807, CTS7121/Z2808,
CTS7267/Z2809, CTS7386/Z2810, CTS7477/Z2811, CTS7652/Z2812,
CTS7751/Z2813, CTS7949/Z2814, CTS8170/Z2816, CTS8224/Z2817,
CTS8394/Z2818, CTS8652/Z2820, CTS8708/Z2821, CTS8716/Z2822,
CTS9010/Z2824, CTS9258/Z2825, CTS9288/Z2826, CTS9368/Z2827,
CTS9405/Z2828, CTS9486/Z2829, CTS9487/Z2830, CTS9518/Z2831,
CTS9648/Z2832, CTS9749/Z2833, CTS9900, CTS10062/Z2834, CTS10140/Z2837,
CTS10338, CTS10613/Z2840, CTS11036/Z2863 to tree on 26 April 2016.
Added CTS11091/Z2865, CTS11126/Z2866, CTS11166/Z2867,
CTS11214/Z2868, CTS11503/Z2869, CTS11526/Z2870, CTS11534/Z2871,
CTS11552/Z2872, CTS11751/Z2874, CTS11775/Z2875, CTS11783/Z2876,
CTS11838/Z2879, CTS11950/Z1880, CTS12008/Z2881,
F3033.2/M1443.2/V4056.2/Z2838.2, FGC2422/Z2715, FGC2425/Z2714,
FGC2427/Z2713, FGC2430/Z2699, FGC2431/Z2710, FGC2432/V1339/Z2712,
FGC2435/Z2698, FGC2436/Y1879, FGC2439/Z2711, FGC2440/V1427/Y3658,
FGC2471/Y1882, FGC2483/Y1933, L495/Z2788 to tree on 27 April 2016.
Added L509/Z2864, L575/V4059/Z2839,
L740/V3959/YSC0000296/Z2836, L750/YSC0000292/Z2725,
L764/YSC0000271/Z2849, L841/YSC0000257/Z2751, L842/YSC0000264/Z2789,
L843/YSC0000273/Z2878, L848/YSC0000299/Z2877, Page123/Z2763,
S22865/Z2845, V1423/Z2717, V1771/Z2724, V2264/Z2728, Y1886,
YSC0000259/Z2761, YSC0000301/Z2882, Z2718, Z2719, Z2721, Z2722, Z2723,
Z2726, Z2727, Z2729, Z2730, Z2731, Z2733, Z2736, Z2737, Z2738, Z2740,
Z2741, Z2742, Z2743, Z2744, Z2747, Z2823, Z2841, Z2842, Z2843, Z2844,
Z2846, Z2847, Z2848, Z2850, Z2851, Z2852, Z2853, Z2854, Z2855, Z2856,
Z2857, Z2861, Z2862, Z2885, Z2887 to tree on 28 April 2016.
Added ZS2286 to tree; removed M1460/PF3816 not meeting
quality guidelines on 30 April 2016.
Added V2740 to tree on 9 May 2016.
Added L300 and L1248 as projects on 28 June 2016.
Added Y18119/Z17925, S2304/Z17937, Z17943, Z41181, Z41182,
Z41183, Z41184, Z41185 on 11 July 2016.