Y-DNA Haplogroup L and its Subclades - 2016

The entire work is identified by the Version Number and date given on the Main Page. Directions for citing the document are given at the bottom of the Main Page.

Version History     Last revision date for this specific page: 15 September 2016

Because of continuing research, the structure of the Y-DNA Haplogroup Tree changes and ISOGG does its best to keep the tree updated with the latest developments in the field. The viewer may observe other versions of the tree on the Web. Email Ray Banks if the differences need clarification or if you find any broken links on this page.

LINKS:  Main Page   Y-DNA Tree Trunk    SNP Index    Papers/Presentations Cited    Glossary   Listing Criteria
COLOR CODES:  Newly confirmed in 2016 within subclade  Confirmed within subclade   Investigational items
^ Indicates a next-generation sequencing entry which does not yet meet quality guidelines for minimum number of reads.
^^ Indicates an entry which does not meet quality guidelines but may be helpful.
~ Indicates only an approximate location on the tree.

The criteria for a representative SNP printed in bold for a subclade is: traditional usage, testing one in multiple labs, and/or being found in the area of the chromosome used in recent research studies.

SNPs listed below in italics (colored black or red) are quality variants from next-generation sequencing reports consistently showing as representing that subgroup.

Contact Person for Haplogroup L: Gareth Henson

L   M20/PF5570 L656, L855, L863, L878/PF5524, L879/PF5697, L1304, M11, M61/Page43, M185/PF5755, M2356/PF5683, M2358, M2362/PF5692, M2365, M2366, M2368/PF5693, M2369/PF5694, M2370/PF5695, M2379/S10007, M2381, M2382/PF5701, M2382/PF5701, M2390/PF5705, M2403/PF5528, M2404/PF5715, M2407/PF5717, M2411/PF5718, M2413/PF5720, M2426/PF5723, M2444/PF5729, M2453/PF5737, M2454/PF5738, M2457, M2463/PF5743, M2464, M2466/PF5744, M2470, M2472/PF5747, M2479, M2482/PF5750, M2485/PF5754, M2489, M2491, M2492/PF5756, M2495, M2497, M2503, M2504, M2518/PF5767, M2522, M2525/PF5771, M2528, M2530, M2531, M2532/PF5773, M2536/PF5775, M2537, M2541/PF5777, M2548, M2551/PF5783, M2554/PF5785, M2556/PF5787, M2559, M2569/PF5792, M2572/PF5795, M2578/PF5805, M2579/PF5806M2583, M2587/PF5810, M2594, M2595/PF5813, M2596, M2597/PF5563, M2598/PF5815, M2607/S7544, M2609, M2611/PF5564, M2612, M2613/PF5821, M2615/PF5823, M2619, M2620, M2621, M2624, M2628, M2629, M2633, M2638, M2640/PF5829, M2642, M2644, M2645, M2650, M2655/PF5832, M2657/PF5833, M2660/PF5837, M2661/PF5839, M2670/PF5841, M2675/PF5842, M8987/PF5699, PF5532, PF5539, PF5554, PF5557, PF5583, PF5732, PF5757,  PF5779, PF5790, PF5791, PF5802, PF5804, PF5830, PF5835, Y5514, Y5529/Z20172, Y5530/Z20173, Y5564/Z20190, Z20117
L~ FGC11566, M2363, M2456, M2474, M2500, M2526/PF5772, M2527, M2552, M2564, M2584/PF5560, M2612, M2637/PF5827, M2646,  M2652, M2680, PF5790, Y5543/Z20166, Y5553/Z20165, Z20055, Z20116, Z20164, Z20192
L1   M22, L1304, M295, Page121
L1~ CTS2285/M2465, M2374, M2383, M2384, M2386, M2389/PF5703, M2391, M2392, M2394, M2395/PF5710, M2399, M2405, M2406, M2409, M2416, M2418, M2419, M2420, M2421, M2427/PF1963/Z1075, M2430/PF5725, M2435/PF5726, M2437, M2438, M2445, M2460, M2462, M2468, M2471, M2490, M2494, M2496, M2517, M2543, M2544, M2545, M2546, M2560, M2568, M2576, M2586, M2593, M2600, M2602, M2614, M2617, M2631, M2635, M2639, M2665,  M2666, M2668/PF5840, M2673, M2676, SK1410/Y5528/Z21069, Y5517/Z20124,  Y5520/Z20128, Y5522/Z20156, Y5523/Z21057, Y5524/Z20161, Y5534/Z20201, Y5537/Z20202, Y5545/Z20129, Y5546/Z21058, Y5555/Z20166, Y6264/Z20194, Z20130, Z21036, Z20138, Z20139, Z20140, Z20141, Z20153, Z20154, Z20155
  L1a   M2481, M2565, M2605, M2664, Y5525/Z20163, Y5560/Z20168
  L1a~ M2357, M2375, M2401, M2480, M2498, M2555, M2623, SK1417/Y5544, SK118/Y5565, Z20132, Z20135, Z20170
   L1a1   M27, F11470/Z20390, M76, P329, L1319, L1320, L1321, SK1423, SK1419/Z20362. SK1420/Z20446, SK1421/Z20361, SK1422/Z20416, Z8046, Z8047, Z8049, Z8050, Z20336, Z20354, Z20355, Z20356, Z20357. Z20358^, Z20359, Z20360, Z20363, Z20464, Z20365, Z20367, Z20368, Z20369^, Z20370, Z20371, Z20372, Z20373^, Z20376, Z20378, Z20379, Z20380, Z20382, Z20383, Z20384, Z20385, Z20387, Z20388, Z20389, Z20391, Z20392, Z20393, Z20394, Z20396, Z20397, Z20398, Z20399. Z20400, Z20401, Z20403, Z20405, Z20406, Z20407, Z20410, Z20411, Z20412, Z20413, Z20414, Z20415, Z20418, Z20419, Z20422, Z20424, Z20425, Z20426, Z20427, Z20428, Z20433, Z20434, Z20435, Z20436, Z20437, Z20438, Z20439, Z20440, Z20441, Z20442, Z20443, Z20444, Z20445, Z20448, Z20449, Z20450, Z20452, Z24638.2
   L1a1~ Z8048, Z20402, Z20429
    L1a1a  Z20458, Z20462, Z20463, Z20464, Z20465, Z20466, Z20468, Z20469, Z20470, Z20472
    L1a1a~ Z20457
     L1a1a1~ Z34426, Z34427, Z34428, Z34429, Z34430
     L1a1a2  Z20475, Z20476, Z20477, Z20483, Z20484, Z20479, Z20480, Z20481, Z20482
     L1a1a2~ Z5925, Z20478
      L1a1a2a~  Z42477, Z42478, Z42479, Z42481
      L1a1a2b~  Z42482, Z42483, Z42484, Z42485, Z42486
    L1a1b  Z5926, SK1426/Z20512, Z20492, Z20493, Z20496, Z20498, Z20499, Z20502, Z20503, Z20504, Z20506, Z20507, Z20508, Z20509
    L1a1b~ Z20497, Z20505
     L1a1b1  Z20519^
     L1a1b1~ M3904
      L1a1b1a~ Z34406. Z34408, Z34409. Z34410
      L1a1b1b~ Z34411. Z34412, Z34413. Z34414, Z34415
     L1a1b2  Z20520, Z20521, Z20522
      L1a1b2a  Z5929
       L1a1b2a1~ Z34379, Z34381, Z34382
       L1a1b2a2~ Z34383, Z34384, Z34385, Z34387
      L1a1b2ab~ Z34467, Z34468, Z34470, Z34471 
     L1a1b3  Z5930
      L1a1b3a  Z5931
       L1a1b3a1  Z5933
        L1a1b3a1a  Z5934
         L1a1b3a1a1  Z20500
          L1a1b3a1a1a~ Z34416, Z34419, Z42487, Z42488 
          L1a1b3a1a1b~ Z34425, Z42489, Z42490, Z42491
         L1a1b3a1a2  Z8033, Z20527, Z20528
          L1a1b3a1a2a  Z5937, Z20535, Z20537, Z20539, Z20540, Z20542, Z20544, Z20545, Z20547, Z20548, Z20550
          L1a1b3a1a12a~ Z20541, Z20546, Z20549, Z20552
           L1a1b3a1a2a1   Z5938, A6402.2/JFS00004.2/Z20559.2, Z8034, Z20558, Z20561, Z20562, Z20563^, Z20564, Z20565
            L1a1b3a1a12a1a~ Z39168.2, Z38449
            L1a1b3a1a12a1b~ Z38450, Z38452, Z38453, Z34113
           L1a1b3a1a12a2~ Z34122, Z34125, Z42492, Z42493
          L1a1b3a1a12b~ Z34479, Z34481, Z34482, Z42494, Z42495
        L1a1b3a1b~  Z34368, Z34369, Z42496, Z42497, Z42498
       L1a1b3a2   Z20568
       L1a1b3a2~ Z5932
        L1a1b3a2a~ Z42499, Z42500, Z42501, Z42502, Z42503
        L1a1b3a2b~ Z34132, Z42504, Z42505, Z42506, Z42507
        L1a1b3a2c~ Z34136, Z42508, Z42509, Z42510, Z42511
      L1a1b3b SK1428, Z34370, Z34371, Z34372
      L1a1b3c FGC18043/SK1433, Y17158
       L1a1b3c1  Z34402, Z34403, Z34404, Z42512, Z42513 
    L1a1c~ F3987
    L1a1d~ CTS12086
   L1a2   M357, L1307^^, M2355^, M2360, M2361, M2364^, M2367^, M2371, M2380, M2393, M2402^, M2412^, M2424^, M2429, M2432, M2439^, M2441^, M2451, M2461^, M2469, M2475^, M2476, M2507, M2523, M2524, M2533^, M2535^, M2540, M2549^, M2557, M2558, M2561^, M2574^, M2575, M2581, M2592^/PR2941^, M2601^, M2606^, M2608^, M2618, M2627, M2630^, M2632^, M2653, M2659, M2662^, Z2662, M5710, SK1446/Y5518, SK1447/Y5559/Z20251, Y5526^/Z20246^, Y5515^/Z20229^, Y5527/Z20248, Y5531^/Z20252^, Y5548^/Z20240^, Y5551/Z20243, Y5552^/Z20244^, Y5554^/Z20247^, Y5556/Z20249, Y5558^/Z20250^, Y5567/Z20256, Y12414^/Z20263^, Z20228^, Z20230^, Z20233^, Z20236, Z20238^, Z20241, Z20262^
   L1a2~ M2387
    L1a2a  M2398, Y12415/Z20266, Z5919
     L1a2a1  Z5920, Y12416/Z20271, Y12417/Z20273, Y12890/Z20274, Z20276
      L1a2a1a  Z20284, Z5921^, Z20282^, Z20283^, Z20285, Z20286^, Z20287^, Z20288^, Z20291^, Z20292^, Z20293^, Z20294^, Z20295^, Z20296^, Z20297, Z20298^, Z20299^, Z20302^, Z20303, Z20304^, Z20305, Z2388.3^
       L1a2a1a1  Z8030, Z20306
        L1a2a1a1a  Z5923, S24402, Z20309, Z20309^, Z20310, Z20311, Z20312, Z20313, Z20314, Z20315, Z20317, Z20319^, Z20320^, Z20321^, Z20322^, Z20323^, Z20324, Z20325^, Z20326, Z20327, Z20328, Z20329, Z20330^, Z20333, Z20335
         L1a2a1a1a1~ Z34393, Z34394, Z34395, Z34396
         L1a2a1a1a2~ Z34397, Z34398, Z34400
        L1a2a1a1b~ Z34475, Z34476, Z34478, Z42525, Z42526
       L1a2a1a2~  Z34373, Z34374, Z34375, Z34376, Z34377
      L1a2a1b~ Y12419, Y17951 
       L1a2a1b1~ CTS1018/M5840, Z34483, Z34484, Z34485, Z34486
       L1a2a1b2~ FGC40759, FGC40771, SK1454, SK1455, Y17950, Y17952, Y17953, Y17954, Y17955, Y17957, Y17958, Y17959, Y17960, Y17961, Y17962, Y17963, Y17965, Y17966, Y17967, Y17968, Y18179, Y18181, Y18182, Y18183, Y18185, Y18186, Y18187, Y18188, Y18189, Y18190, Y18191, Y18193, Y18194, Y18195, Y18196, Y18197, Y18198, Y20047
        L1a2a1b2a~ Y17956, Y18180, Y18178, Y18184, Y18192, Y20048
     L1a2a2~  Z34463, Z34465, Z34466, Z42523, Z42524
    L1a2b~ FGC32285/Y6288
     L1a2b1~ FGC32252/Y6250, FGC32253/Y6251, FGC32255/Y6252, FGC32257/Y6270, FGC32262/Y6256, FGC32264/Y6272, FGC32265/Y6273, FGC32268/Y6274, FGC32269/Y6276, FGC32270/Y6277, FGC32272/Y6279, FGC32276/Y6259, FGC32279/Y6282, FGC32280/Y6283, FGC32281/Y6260, FGC32282/Y6284, FGC32284/Y6286, FGC32287/Y6289, FGC32292/Y6291, FGC32294/Y6293,  FGC32296/Y6295, FGC32297/Y6296, PH2254/Y11220, Y6285, Y13388, Y19628, Y20208
      L1a2b1a~ FGC32248, FGC32250, FGC32254, FGC32256, FGC32260, FGC32263, FGC32266, FGC32267, FGC32271, FGC32273, FGC32274, FGC32278, FGC32283, FGC32286, FGC32288, FGC32291, FGC32293, FGC32295, Y20686, Z6953
      L1a2b1b~ FGC23830/Y6287, Y6246, Y6247, Y6253, Y6254, Y6255, Y6261, Y6265, Y6266, Y6267, Y6268, Y6269, Y6271, Y6275, Y6278, Y6281, Y6292, Y6294, Y19630, Y20206
       L1a2b1b1~ Y6257, Y6258, Y6268, Y6290, Y19627
        L1a2b1b1a~ Y19629, Y19631
    L1a2c~ F2419.2/PF3400.2
  L1b   FGC36877/M317, L655, PH982, PH1071, PH1379, PH2312, PH2342, PH3435, PH3443, PH3615, PH3912, PH4035, PH4159, PH4188, SK1413
   L1b1   M349, Page113
    L1b1a   Page116
    L1b1b~ B374, Z34118, Z34120, Z34121
    L1b1c~ B373, Z34114, Z34115, Z34116, Z42527, Z42528
    L1b1d~ AM00711.2/F2544.2
   L1b2   SK1412
    L1b2a~ M274, SK1415, SK1416
    L1b2b~ PH8, PH108, PH612, PH997, PH1060, PH1371, PH1428, PH1520, PH1844, PH2124, PH2161, PH2832, PH2952, PH3298, PH3524, PH4160, PH4366.1, PH4489, PH4587
     L1b2b1~ PH2079
      L1b2b1a~ PH1714, PH3523
     L1b2b2~ PH1099, PH3342
      L1b2b2a~ PH1728
      L1b2b2b~ Y18889, Y18890
    L1b2c~ SK1414
   L1b3~ CTS7154.2/M5226.2/PF1724.2
    L1b3a~ CTS3080/S1565/V2529
L2   L595, L877/PF5780, Page6/PF5751
 L2~ L865, L868, L869, L870, PF5533, PF5538, PF5540, PF5551, PF5553, PF5569, PF5573, PF5577, PF5579, PF5585, PF5682, PF5684, PF5685, PF5691, PF5696, PF5698, PF5700, PF5706, PF5708, PF5711, PF5713/M11114/V1591, PF5716, PF5721, PF5722, PF5730, PF5731, PF5739, PF5740/Z20586, PF5741, PF5742, PF5746, PF5748, PF5749, PF5752, PF5753, PF5758, PF5759, PF5760, PF5761, PF5762, PF5763, PF5764, PF5765/S7336, PF5766, PF5768, PF5769, PF5770, PF5774, PF5776, PF5778, PF5782, PF5784, PF5786, PF5788, PF5789, PF5796, PF5797, PF5798, PF5799, PF5800, PF5803, PF5807, PF5808, PF5809, PF5811, PF5812, PF5817, PF5818, PF5819, PF5820, PF5822, PF5824, PF5825, PF5826, PF5828, PF5831, PF5834, PF5836, PF5838, PF5843, Z991, Z20571, Z20572, Z20573, Z20574, Z20575, Z20576, Z20577, Z20578, Z20579, Z20581, Z20582, Z20583, Z20584, Z20585, Z20587, Z20588, Z20589, Z20590, Z20591, Z20592, Z20593, Z20594, Z20595, Z20596, Z20597, Z20598, Z20599, Z20600, Z20601, Z20602, Z20603, Z20604, Z20605, Z20606, Z20607, Z20608, Z20609, Z20610, Z20611/YP914, Z20612, Z20613, Z20614, Z20615, Z20616, Z20617, Z20618, Z20619, Z20620, Z20621, Z20622, Z20623, Z20624, Z20625, Z20626, Z20627, Z20628, Z20629, Z20630, Z20631, Z20632, Z20635, Z20636, Z20637, Z20638, Z20639, Z20640, Z20641, Z20643, Z20644, Z20645

Y-DNA haplogroup L is found at low frequencies in the Middle East and Europe as paragroup L-M317 (L1b*, originally L2*), subgroup L-M349 (L1b1, originally L2a) and subgroup L-L595 (L2), and at significant frequencies in South Asia as subgroups L-M27 (L1a1, formerly L1a and originally L1) and L-M357 (L1a2, formerly L1c and originally L3).


Geographic distribution so far of ancestry of modern-day haplogroup L subgroup men (not complete):

L1a1  M2481                    Pakistan, India, Sri Lanka, Bangladesh, Iraq, Bahrain, Kuwait, Saudi Arabia,
                                                 Qatar,United Arab Em.
L1a2  M357                      India, Sri Lanka, Bangladesh, Iraq, Kuwait, Saudi Arabia, Qatar, Armenia, Russia, Italy
L1b    FGC36877/M317   Israel (Druze), Lebanon,  Kuwait, Kazakhstan (Tatars), Armenia, Azerbaijan, Iran, 
                                                 Turkey, Belarus, Greece, Bulgaria, Germany, Austria, Switzerland, Portugal,
                                                 Ukraine, Russia, Mongolia
L2      L595                       Germany, Sardinian Italy


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A limited number of qualifying investigative items here appear only in the public Y trees of Family Tree DNA and/or YFull.

Additional Resources:
ISOGG Wiki - What you need to know about Genetic Genealogy.
The Y-Haplogroup L Project, Gareth Henson, Peter Hrechdakian.

Corrections/Additions made since 1 January 2016:

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