Y-DNA Haplogroup O and its Subclades - 2017

The entire work is identified by the Version Number and date given on the Main Page. Directions for citing the document are given at the bottom of the Main Page.

Version History     Last revision date for this specific page: 21 September 2017

Because of continuing research, the structure of the Y-DNA Haplogroup Tree changes and ISOGG does its best to keep the tree updated with the latest developments in the field. The viewer may observe other versions of the tree on the Web. Email Ray Banks if the differences need clarification or if you find any broken links on this page.

LINKS:  Main Page   Y-DNA Tree Trunk    SNP Index    Papers/Presentations Cited    Glossary   Listing Criteria
FONT COLORS:  Newly confirmed in 2017 within subclade  Confirmed within subclade  Investigational items 

^ Indicates a next-generation sequencing entry which does not yet meet quality guidelines for minimum number of reads.
^^ Indicates an entry which does not meet quality guidelines but may be helpful.
~ Indicates only an approximate location on the tree.

The criteria for a representative SNP printed in bold for a subclade is: traditional usage, testing one in multiple labs, and/or being found in the area of the chromosome used in recent research studies.

SNPs listed below in italics (colored black or red) are quality variants from next-generation sequencing reports consistently showing as representing that subgroup.

Contact Person for Haplogroup O: Owen Lu 

O   M175, CTS647, CTS874/M1530, CTS1182/M1740, CTS1658/M1746, CTS1853/M1747, CTS2340, CTS2419/M1748, CTS3283/M1749, CTS3771/M1750, CTS5118/M1752, CTS5493/M1753, CTS7498/M1758, CTS7553/M1759, CTS8384/M1762, CTS8746/M1763, CTS9039/M1766, CTS9544/M1767, CTS9577, CTS9602/M1768, CTS10784/M1779, CTS10995/M1780, CTS11955/M1781, F14/M1738, F45/M1739, F84/M1741, F137/M1743, F171/M1573, F175, F182/M1744, F282/M1359, F315/M1751, F355/M1754, F371/M1756, F380/M1757/S27659, F452/M1761, F478/M1764, F494/M1765, F537/M1769, F540, F546/M1770, F600/M1777, F614/M1778, F668/M1782, FGC12433, FGC12434, FCG12436, L1361/M1771, M1742, M1745, M1772, M1773, M1774, M1775, M1776, P186, P188, P191, P196, Z16962, Z16966
O1   F265/M1354, CTS2866, F75/M1297, F429/M1415, F465/M1422
� � O1a   M119, CTS31, F589/Page20, L246, L466
� � � O1a1   B384/Z23193
� � � � O1a1a   M307.1/P203.1, CTS3422, CTS4351, CTS5059,CTS6864, CTS7015, CTS8229, CTS8875, CTS8934, CTS9321, CTS11688, F31, F54, F89, F303, F333, L83
� � � � O1a1a1   F446, CTS4588, F560, FGC15381/K620/Z23389, K613/Z23387, K619/Z23388
� � � � O1a1a1a   F140, CTS611, CTS3089, CTS3265, CTS3269,CTS11270, F157, F343, F424, F518, F571, FGC15382/Z23466, FGC15391/Y14275/Z23470, FGC15383/K625/Z23474, V68.2, Z23457
� � � � � � O1a1a1a1   F78CTS1711, CTS2268, CTS3758, CTS4478, CTS9282, CTS10963, CTS11692, F168, F206,  FGC15386/Z23467, FGC15388/K642, FGC15394/K618/Z23468, K614/Z23459, Y23446, Z23458, Z23471
� � � �  � � � O1a1a1a1a   F81, CTS4910, CTS5709, FGC15392/Y14265/Z23469
� � � �  � � � � O1a1a1a1a1  CTS2458
� � � �  � � � �  O1a1a1a1a1a   F533
� � � � � � � �  O1a1a1a1a1a1   F492, CTS2498, CTS2594, CTS4206,CTS5075, CTS11784, F619, K629/Z23478
� � � � � � � �   O1a1a1a1a1a1a   F656, Z23481
� � � � � � � �   O1a1a1a1a1a1a1  A12440
� � � � � � � �   O1a1a1a1a1a1a1a  A12439 
� � � � � � � �   O1a1a1a1a1a1a2  A14788, A14789  
� � � � � � � �   O1a1a1a1a1a1a3  F65, F285, F469
� � � � � � � �   O1a1a1a1a1a1a4  MF1068, MF1069, MF1070
� � � � � � � �   O1a1a1a1a1a1a5  Z23482, Z23484, Z23485, Z23486, Z23487, Z23488, Z23489
� � � � � � � �   O1a1a1a1a1a1b   FGC66168, Z23505
� � � � � � � �   O1a1a1a1a1a1b1 CTS11553, Z23494, Z23496, Z23503 
� � � � � � � �   O1a1a1a1a1a1c   Y31266, Y31267
� � � � � � � �   O1a1a1a1a1a1c1   Y31261, Y31262, Y31263, Y31264, Y31265, Y31643
� � � � � � � �   O1a1a1a1a1a1d   A12441, A12442, A12443
� � � � � � � �   O1a1a1a1a1a1e   MF1071, MF1072, MF1073
� � � � � � � �   O1a1a1a1a1a1e1   MF1074
� � � �  � � � �  O1a1a1a1a1a2   CTS4585, CTS7624, FGC15395/Y14269, FGC15397/Y14272, FGC15398/Y13987, FGC15399/Y13986, FGC15400/Y13989, FGC15401/Y13988, FGC15402/Y13984, FGC15409/Y13985, FGC47328, FGC47336, Y14273 
� � � �  � � � � O1a1a1a1a2  MF1075, MF1076, MF1077, MF1078, MF1079, MF1080, MF1081, MF1082
� � � � � � O1a1a1a2   YP4610/Z39229
� � � � � � � O1a1a1a2a   AM00330/AMM480/B386, SK1533, Z39230/YP4600, Z39231/YP4601, Z39232/YP4603, Z39233/YP4605, Z39234/YP4607, Z39235/YP4609, Z39236/YP4611, Z39237/YP4612, Z39238/YP4613, Z39239/YP4614, Z39240/YP4615, Z39241/YP4616, Z39242/YP4617, Z39243/YP4618, Z39244/YP4619, Z39245/YP4620, Z39246/YP4621
� � � � � � � � O1a1a1a2a1   AM00333/AMM483/B387, Z39247/YP4602
� � � �  � � � � O1a1a1a2a1a   B388, Z39248/YP4604, Z39249/YP4608
� � � �  � � � O1a1a1a2b   SK1555
� � � � � O1a1a1b   SK1568/Z23420
� � � � � � O1a1a1b1   M101
� � � � � � O1a1a1b2   Z23392, Z23404, Z23440
� � � � � � � O1a1a1b2a   Z23442
� � � � � � �  �O1a1a1b2a1   SK1571, Z39268
� � � �  O1a1a2   CTS52, CTS5880, CTS8423, CTS11785, F4084
� � � �  � O1a1a2a   CTS701, CTS4115, CTS6217
� � � � � � O1a1a2a1   K644/Z23266, Z23269
� � �  �O1a1b   CTS5726, CTS3085, CTS3400, CTS7370^,CTS8228^, CTS9138^, CTS9243^, CTS9540^, CTS11079, F819^, F1452, F1620.2, F2104, F2644^, F2852^, F3063, F3495,F3640, SK1589, Z39164, Z39165, Z39166^, Z39167, Z39168, Z39169, Z39170, Z39171^, Z39172^, Z39173, Z39174,Z39175^, Z39176, Z39177, Z39178, Z39179^, Z39180^, Z39181, Z39182, Z39183^, Z39184, Z39185, Z39186, Z39187,Z39188, Z39189, Z39190, Z39191, Z39192, Z39193, Z39194, Z39195, Z39196^, Z39197, Z39198^, Z39199^
� � � � O1a1b~ F2911
� � � O1a2   M110, F881, F908, F1081,F1608, F1880, F2340, F2380, F2635, F2671, F3134, F3152, F3157, F3214, M50, M103
� � � � O1a2a   F3288
� � � � O1a2a1   B392, Z38606,Z38607, Z38608, Z38609, Z38610, Z38611
� � � � � O1a2a1a   B393, Z38612
� � � O1a3~ Page109
� � O1b   M268, F138, F167/L690, F256/M1341, F292/M1363, F330/L463, F435/M1417, F516, P31
� � � O1b1   F2320, F3008/M1441, F3286/M1465
� � � � O1b1a   M1470
� � � � O1b1a1   PK4, CTS6616/M1392, CTS7320/M1405, CTS7541/M1407, CTS7548/M1408, CTS8589/M1425, CTS9150, CTS11303, F722/M1279, F932/M1290, F1058/M1302, F1085/M1305, F1130/M1306, F1265.1/M1313.1, F1346/M1316, F1353/M1317, F1809/M1352, F2317, F2338.2/M1391.2, F3033.1/M1443.1/V4056.1/Z2838.1, F3424/M1489, F3439/M1494
� � � � � O1b1a1a   M95, F709, F987/M1300, F1931/M1362, F2176/M1376, F2254/M1383, F2917/M1436, F4233/M1404
� � � �  � � O1b1a1a1   F1803/M1348, CTS6251/M1384, M1310
� � � �  � � �  O1b1a1a1a   F1252
� � � �  � � � � O1b1a1a1a1   F2924, CTS10077, CTS10240
� � � � � � � � �  O1b1a1a1a1a   M111, M88, CTS1163, F1120, F1643, F2346, F2454, F2524, F2554, F2861, F3055
� � � � � � � �  � O1b1a1a1a1a1   F2758, CTS10965, Z24081, Z24082, Z24084, Z24085, Z24087
� � � � � � � �  � � O1b1a1a1a1a1a   Z24083, Z24086
� � � �  � � � �  � � � O1b1a1a1a1a1a1   Z24089
� � � �  � � � � � � � � O1b1a1a1a1a1a1a   F923, F1108, F2411, F3671
� � � �  � � � � � � � � O1b1a1a1a1a1a1a1   CTS2022
� � � �  � � � �  � � � � � O1b1a1a1a1a1a1a1a   F1399, F3053
� � � �  � � � � � � � � � � O1b1a1a1a1a1a1a1a1   F2415, CTS3745, Z3955.1
� � � �  � � � � � � � � O1b1a1a1a1a1a1a2   Z24131
� � � � � � � �  � � � � O1b1a1a1a1a1a1a3   Z24100
� � � � � � � � � � � � O1b1a1a1a1a1a1b   SK1627/Z24091, Z24093
� � � �  � � � � � � � � O1b1a1a1a1a1a1b1   Z39410
� � � � � � � �  � � � O1b1a1a1a1a1a2   Z24088
� � � �  � � � �  � O1b1a1a1a1a2   F2890
� � � � � � � �  � � O1b1a1a1a1a2a   Z24048
� � � � � � � �  � � � O1b1a1a1a1a2a1   Z24050
� � � �  � � � � � � O1b1a1a1a1a2b   Z24014
� � � � � � � � O1b1a1a1a1b   CTS5854, Z23743, Z23744
� � � � � � � � � O1b1a1a1a1b1   Z23810
� � � � � � � �  � � O1b1a1a1a1b1a   CTS7399, FGC19685, FGC19699 FGC19670/Y14253/Z23825, FGC19672/Y14260/Z23840, FGC19673/SK1620/Y14000/Z23821, FGC19674/Y14018/Z23835, FGC19675/Y13998/Z23819, FGC19676/Y14255/Z23831, FGC19677/Y14259/Z23834, FGC19678/Y14257, FGC19679/Y14008/Z23828, FGC19681/Y14020/Z23836, FGC19682/Y14261/Z23841, FGC19683/Y14011/Z23829, FGC19684/Y14014/Z23832, FGC19686/Y14022/Z23839, FGC19687/Y13999/Z23820, FGC19688/Y14007/Z23827, FGC19689/SK1619/Y14001/Z23822, FGC19694/SK1618/Y14256, FGC19705/Y14010, FGC19710/Y14021/Z23837, Z23824, Z23833, Z23838, Z23843
� � � � � � � �  � � � O1b1a1a1a1b1a1   FGC19713/Y14026, FGC19671/Z23845, FGC19690/Y14251/Z23851, FGC19693/Y14013/Z23857, FGC19698/Y13996/Z23852, FGC19700/Y14017/Z23861, FGC19701/Y14002/Z23853, FGC19703/Y14262/Z23866, FGC19711/Y14025/Z23865, FGC19712/Y14016/Z23860, FGC19715/Y13997, FGC19724, Z23858
� � � � � � � �  � � � � O1b1a1a1a1b1a1a   Z23849
� � � � � � � �  � � � �  O1b1a1a1a1b1a1a1   FGC61038, FGC61039, FGC61054, Y31276, Y31927, Y31928  
� � � �  � � � � � � O1b1a1a1a1b1b   CTS651, CTS5267, CTS7642, CTS8722, CTS9915
� � � � � � � � � � � O1b1a1a1a1b1b1   CTS9884
� � � � � � � � � O1b1a1a1a1b2   F4229, F4212, F4251, FGC61079, FGC61080, PF3452, Z23746, Z23769, Z23781, Z23787, Z23799   
� � � � � � � �   O1b1a1a1a1b2a   F809, Y31259, Y32109, Z23745, Z23766, Z23777, Z23784, Z23786, Z23791, Z23797, Z23801, Z23803, Z23804, Z23807
� � � � � � � �   O1b1a1a1a1b2a1   F2517, Z23795
� � � �  � � � � O1b1a1a1a2   SK1630, SK1631, SK1632, SK1633
� � � � � � � � O1b1a1a1a2a   SK1636
� � � � � �  �O1b1a1a1b   F789/M1283, F2028/M1367,F2522/M1410, F3092, F3318/M1469, F4181/M1357
� � � � � � � � O1b1a1a1b1   FGC29900/Y9322/Z23667
� � � � � � � � O1b1a1a1b1a   B426/FGC29896/Y9033/Z23671, FGC29897/Y9320/Z23673, FGC29898/Y9036/Z23674, FGC29899/Y9034/Z23672
� � � � � � � �  � O1b1a1a1b1a1   FGC29907/YP3930, SK1640/YP3931
� � � � � � � � � O1b1a1a1b1a2   B427/Z23680, M6661
� � � � � � � � O1b1a1a1b1b   Z39485 , Z39486^, Z39487, Z39488, Z39489^
� � � � � � � � O1b1a1a1b1c   B418, Z39490
� � � � � � � � O1b1a1a1b2   SK1646
� � � �  � � O1b1a1a2   CTS350, CTS2549, CTS3471,CTS6531, CTS7187, CTS7270, CTS7809, CTS11962.2/Z2953.2
� � � �  � � O1b1a1a3~ Page103
� � � � � O1b1a1b   F838, CTS892
� � � � � � O1b1a1b1   F1199, F1309, F1685,F1742, F2189, F2445, F2995, F3353
� � � � O1b1a2   Page59, CTS1456, CTS1719, CTS4040, CTS5528, CTS5745, CTS7643, F883, F888, F1098, F1485, F2117, F2290, F2601, F3016, F3079, F3445
� � � � � O1b1a2a   F993, CTS2335, CTS5445,CTS7866, CTS8849, CTS10154, CTS10437, CTS11033, F1484, F1669, F2120, F2124, F2131, F2383, F2414, F2590, F2771,F2862, F3036, F3135, F3217, F3262, SK1659, Z24391
� � � � � � O1b1a2a1   F1759, CTS5847,CTS8414, CTS11890, F2064, F3314, F3323, F3346, F3478, Z24393, Z24407
� � � �  � � � O1b1a2a1a   CTS1127, A16611, A16612, A16613, CTS6105, PH2506, PH2797, PH5023
� � � � � O1b1a2b   F417/M1654, CTS7268, CTS10113,F1236, F1860, F1956, F2061, F2504, F2775, F3252, F3291, F3632
� � � � � � O1b1a2b1   F840, CTS250, CTS949, F984, FGC30073, Y10774
� � � �  � � �O1b1a2b1a   F1127, CTS10685, F1628, F1776, F2183
� � � �  � � O1b1a2b2   CTS1451, CTS4475, CTS5284, CTS6503, CTS9341
� � � � � O1b1a2c   CTS9996
� � � O1b2   P49, M176/Page63/SRY465
� � � � O1b2a   F1942/Page92, IMS-JST022454, M302
� � � � O1b2a1   CTS9259, CTS348, CTS9752, F2315, PF4474, Z24512, Z24520, Z24522, Z24553
� � � �  � O1b2a1a   F1204, F1290, F1530, F1805,F2634, F2960, F3147, F3327, F3356, K10
� � � � � � O1b2a1a1   CTS713, CTS7776, CTS8032, CTS10145, CTS10341, CTS10674, CTS11723, CTS11986, K7
� � � �  � � � O1b2a1a1a   CTS1875, CTS7465, CTS8777
� � � �  � � � � O1b2a1a1a1   CTS10682, CTS1966,CTS3764.2, CTS3926, CTS6725, CTS7366, CTS8034, Z24596
� � � � � � � O1b2a1a1b   Z24598, Z24599
� � � �  � � � O1b2a1a1c   CTS203, CTS6622, CTS9892
� � � �  � � O1b2a1a2   F2868, F3110, K4
� � � � � � � O1b2a1a2a   L682, CTS2734, CTS6803,CTS9322, K28
� � � � � � � � O1b2a1a2a1   CTS723, Z24624
� � � �  � � � � O1b2a1a2a1a   CTS7620, Z39506
� � � �  � � � � O1b2a1a2a1b   A12446, A12447, A12448
� � � �  � � � �  � O1b2a1a2a1b1   PH40, PH437
� � � � � � � O1b2a1a2b   F940, F1912, F3390
� � � �  � � O1b2a1a3   CTS10687, F1800, F1813, F1823, F1877, F3821
� � � � � � � O1b2a1a3a   CTS1215, CTS3729, F787, F867, F1553, F2026, F2604, F2767, F3065
� � � � O1b2a1b   CTS562, CTS3505, CTS10354
� � � � � O1b2a2~ Page90
O2    M122, CTS10736, CTS10753, F36, F400, F433, F471, F633,P198
� � O2a   M324, CTS8153, CTS8399, F27, F71, F113, F129, F166, F341, F406, F572, P93/Page79, P197, P199, P200
� � � O2a1   L127.1, CTS5687, CTS7638,CTS11981, F51, F58, F95, F145, F159, F164, F170, F243, F272, F300, F304, F305, F412, F498, F573, F601, F642, F676, KL1/L465, KL2/L467
� � � � O2a1a   F1876/Page127, F964, F3143
� � � �  O2a1a1   F2159, F2726, F2789, F3169, F4102
� � � �  � O2a1a1a   F1867/Page124
� � � �  � � O2a1a1a1   F852, CTS3709, CTS8168, CTS8587, CTS10730, F1054, F4348/Z24834, SK1669/Z24838, Z24836, Z24840, Z24844
� � � �  � � O2a1a1a1a   F2266
� � � �  � � � O2a1a1a1a1   L599, M121
� � � �  � � �  O2a1a1a1a1a  Z43961, Z43962
� � � �  � � �  O2a1a1a1a1a1  Z43963, Z43964, Z43965
� � � �  � � O2a1a1a1b   F854
� � � �  �  O2a1a1a1b1   Z43966, Z43967, Z43968, Z43969, Z43970, Z43971, Z43972, Z43973, Z43974
� � � �  �  O2a1a1a1c~ Page130
� � � �  O2a1a1b   F915, CTS7667,  F1675, F2528, F2633, F2975, F3182, F3287, F3417, F3669
� � � �   O2a1a1b1   F1478, F1920, F3046, F3284, F3656, F4195, F4312
� � � �  �  O2a1a1b1a   PF5390
� � � �  �   O2a1a1b1a1  CTS1936, Z24782, Z24797, Z24818
� � � �  �    O2a1a1b1a1a  Z43975, Z43976, Z43977, Z43978, Z43979, Z43980
� � � �  �   O2a1a1b1a2  FGC33994, A18026, FGC34004, FGC34008, FGC34009, FGC34019, FGC34022, FGC34023, FGC34029
� � � � O2a1b~ M164
� � � � O2a1c   IMS-JST002611, CTS1872, CTS2483, CTS5879, F61, F99, F240, F247, F302, F309, F460, F562
� � � � O2a1c1   F18, CTS11342
� � � �  O2a1c1a   FGC12511
� � � �  O2a1c1a1   F117, CTS5820, F514, FGC12471/Y3268
� � � �  O2a1c1a1a   CTS4598, CTS12658, F108, F197, F253, Y3547/Z25052
� � � �  O2a1c1a1a1   CTS4263, CTS4820, F2, F13, F30, F62, F68, F86, F189, F251, F379, F530, FGC12469/Y3286/Z25051, FGC12470/Y3292
� � � �  � �   O2a1c1a1a1a   F11, F60
� � � �  � �   O2a1c1a1a1a1   F325, F362, F425/Page69, F526, F561FGC12472/Y3271/Z25059  
� � � �  � �   O2a1c1a1a1a1a   F632, CTS4580, Z25061
� � � �  � �   O2a1c1a1a1a1a1   F110/M11115, CTS10716, F133, F434, F646, Y21437, Y21448, Z25064, Z25065, Z25067, Z25068  
� � � �  � �    O2a1c1a1a1a1a1a   CTS3663, Z25081
� � � �  � � � �  � �   O2a1c1a1a1a1a1a1   F17, CTS362, CTS2801, CTS4414, CTS5822, CTS6777, CTS9127, CTS11839, F64, F92, F357, F529, F620, F672, F691, Z25075, Z25076, Z25077, Z25079, Z43887
� � � �  � � � �  � �    O2a1c1a1a1a1a1a1a  F377
� � � �  � � � �  � �     O2a1c1a1a1a1a1a1a1  CTS7789, F1249, F1611, F2270  
� � � �  � � � �  � �      O2a1c1a1a1a1a1a1a1a  F856, F1095, F1250, F2427, F2732
� � � � � � � �  � � �  � � � O2a1c1a1a1a1a1a1a1a1   F1418, F1495
� � � � � � � �  � � �  � � � O2a1c1a1a1a1a1a1a1a2   Z25097, Z25098
� � � �  � � � �  � �     O2a1c1a1a1a1a1a1a2  CTS7501
� � � �  � � � �  � �    O2a1c1a1a1a1a1a1b  F793, F1316C, F2035, F2108, F3433, Z43869, Z43870, Z43871, Z43872, Z43873, Z43874, Z43875, Z43876, Z43877, Z43878  
� � � �  � �    O2a1c1a1a1a1a1b    Y20951, Y21440, Y21441Z43879
� � � �  � �     O2a1c1a1a1a1a1b1   Y20939, Y21435, Y21442, Y21443
� � � �  � �      O2a1c1a1a1a1a1b1a    Y20932, Y20933, Y20934, Y20936, Y20938, Y20940, Y20941, Y20942, Y20943, Y20945, Y20946, Y20947, Y20949,  Y21444, Y21446, Y21447, Z34692
� � � �  � �      O2a1c1a1a1a1a1b1a~  Y20935, Y20937, Y20944, Y20948, Y20950, Y21438, Y21439, Y21445, Z43880. Z43881, Z43882, Z43883, Z43884, Z43885, Z43886   
� � � �  � �   O2a1c1a1a1a1b   F38, F106, F142, F178, F286, F381, F386, F475, F479, F485, F553, F592, F625, F3453    
� � � �  � �   O2a1c1a1a1a1b1   F136, F358, F1978     
� � � �  � �   O2a1c1a1a1a1b1a   F270, F3131   
� � � �  � �   O2a1c1a1a1a1c    F12, F196, F480, F659, F660    
� � � �  � �   O2a1c1a1a1a1c1   F683, Z25153
� � � �  � �   O2a1c1a1a1a1d    F930, F1232, F2356, F2589
� � � �  � �   O2a1c1a1a1a1d1   F2685
� � � �  � �   O2a1c1a1a1a1e   F1365/M5420/PF1558, F921, F1676, F2109, F2180, F2213, F3232, F3452, F4062, FGC54466, FGC54513, FGC54523, FGC54527, Y15975, Y15977, Y15978, Y15979,  Y16155
� � � �  � �   O2a1c1a1a1a1e1   Y15976, Y15980, Y26385
� � � �  � �   O2a1c1a1a1a1e1a  Y16154  
� � � �  � �    O2a1c1a1a1a1e1a1  Y26383, Y26387, Y27899
� � � �  � �    O2a1c1a1a1a1e1a1a   SK1686, FGC49730, Y26384, Y26386, Y27894, Y27895, Y27896, Y27897, Y27898, Y27900
� � � �  � �   O2a1c1a1a1a1e2    FGC54486, FGC54495, FGC54520, FGC54522, FGC54529, FGC54538, FGC54540, FGC54545, FGC54488
� � � �  � �   O2a1c1a1a1a1e2a   FGC54507
� � � �  � �   O2a1c1a1a1a1e2b   MF1083
� � � �  � �   O2a1c1a1a1a1f  CTS12877, Z25165
� � � �  � �   O2a1c1a1a1a1f1   F2527
� � � �  � �   O2a1c1a1a1a1f1a   CTS5409, CTS6236  
� � � �  � �    O2a1c1a1a1a1f1a1   CTS9711, MF1308
� � � �  � �    O2a1c1a1a1a1f1a1a   MF1309
� � � �  � �   O2a1c1a1a1a1f1b   F2941, F4073, F4119
� � � �  � �   O2a1c1a1a1a1g   F723, F971, F972, F1210, F1351, F1422, F1638, F2357, F2560, F2719, F3042, F3103, F3270, F4171
� � � �  � �   O2a1c1a1a1a1h   CTS2154, Y29851
� � � �  � �   O2a1c1a1a1a1h1  CTS2107, CTS2128, CTS2916, CTS3518, CTS4150, CTS8491, CTS10670
� � � �  � �   O2a1c1a1a1a1i   SK1691, SK1692, SK1695, SK1696
� � � �  � �   O2a1c1a1a1b    PH203, FGC1803.2/Y2044.2, PH278, PH975, PH1787, PH2385, PH3699, PH3838, PH3849, PH4404, PH4535, PH4997, PH5031, PH5323
� � � �  O2a1c1a2     F449
� � � �  � � � O2a1c1a2a   F238
� � � �  � � � � O2a1c1a2a1   F134, F271, F322
� � � � � � � �  O2a1c1a2a1a   CTS10846, CTS11423, F1894, F2468, F3031, F3245, F16325    
� � � � � � � �  � O2a1c1a2a1a1   CTS6279, F1273, F3256, F3309, F3434, F20597, F25545, F25621
� � � � � � � �  O2a1c1a2a1b   F724, F1156, F1336, F1910
� � � �  � � � O2a1c1a2b   F1266, F1497, F2016, F4267
� � � �  O2a1c1b   CTS498, CTS957, CTS1028, CTS2398, CTS4941, CTS5926, CTS7341, CTS8263, CTS9516, CTS10139, CTS10185, CTS11076, CTS11949
� � � � O2a1c2   FGC3750/SK1673, L133.2/Page74.2
� � � O2a2   IMS-JST021354/P201, CTS8236, F525
� � � � O2a2a   M188
� � � � �  O2a2a1   F2588, CTS3106, F2638
� � � � O2a2a1a   CTS445 , CTS11727, F2055, F2221, F2329, F2592, F3559
� � � �  � � O2a2a1a1   CTS201
� � � �  � � � � O2a2a1a1a   M159/Page96
� � � �  � � O2a2a1a2   M7, CTS10944, CTS12052, F2591, F2594, F2680, F3177, F3342, F3371, F4066, M209, Z25245
� � � �  � � � O2a2a1a2a   F1276, F1898, F2179, F2428, F2962, F3406, F4064, F4173
� � � �  � � � � O2a2a1a2a1   CTS6489
� � � �  � � � � O2a2a1a2a1a   F1275, F2309, F3085
� � � � � � � � O2a2a1a2a1a1   M113
� � � �  � � � � O2a2a1a2a1a2   N5, F1100, F1234, F1411, F3229, F3516
� � � � � � � � O2a2a1a2a1a3   Z25400
� � � � � � � � O2a2a1a2a2   F1863, F2079
� � � �  � � � � O2a2a1a2a2a   F1134, F1992, F2332, F2480, F2612
� � � � � � � � O2a2a1a2a2a1   F1262, F2208, F2515, F3469
� � � �  � � � O2a2a1a2b   Y26403, Y26409, Y26412, Y26413, Y26415, Y26416, Y26421, Y26422, Y26424, Y26428, Y26430, Y26433, Y26434, Y26437, Y26440, Y26446, Y26448, Y26457, Y26460, Y27924, Y27927, Y27932, Y27933, Y27934, Y27939, Y27942, Y27943, Y27951, ZS2079
� � � � O2a2a1b   F1837 , F2576, F2928, F4088, F4163
� � � � O2a2a2   F879
� � � � O2a2a2a   F1226
� � � � � � O2a2a2a1   F2859, F2937
� � � � O2a2b   P164, CTS4723, CTS11109, CTS12099, F130, F131, F299, F422, F427
� � � � O2a2b1   M134, F100/M1546
� � � � O2a2b1a   F450/M1667
� � � �  � � O2a2b1a1   M117, Page23, CTS899/M1531, CTS1275/M1536, CTS3251, CTS5128/M1619, CTS6623/M1638, CTS11742/M1720, F141/M1564, F144, F235/M1587, F342/M1627, F373/M1636, F476/M1671, F579/M1692, F581, F584, F613/M1702, F649
� � � �  � � � O2a2b1a1a   M133, F8, F42
� � � �  � � � � O2a2b1a1a1   F438
� � � �  � � � � O2a2b1a1a1a   Y17728
� � � �  � � � � O2a2b1a1a1a1   F155, CTS630, CTS1304, CTS2760, CTS11316, F201, F254, F276, F316, F414, F440, F541, F703, F6227, F6517, F6530, F6872, F6909^, F7446, F7447, F7556, F7973, F7994, F14281, F14428, F14445, F14663^, Y21365, Y21366, Y21367
� � � �  � � � �  � � O2a2b1a1a1a1a   F813/M6539, Y21364
� � � �  � � � �  � � � O2a2b1a1a1a1a1   Y20928
� � � � � � � � O2a2b1a1a1a2   F1754
� � � � � � � �  � � O2a2b1a1a1a2a   F2137, A16631, A16632, F2503, F2643, S24099.2
� � � � � � � �  � � O2a2b1a1a1a2a1   F1442, F2779
� � � � � � � �  � � O2a2b1a1a1a2a1a   F1123, F2265, F2425, F2453, F3048, F3115
� � � � � � � �  � � � �  O2a2b1a1a1a2a1a1~ F1369
� � � � � � � �  � � O2a2b1a1a1a2a2   A16636, A16637, A16638, A16639, A16640, A16641, A16642, A16643, A16644, A16645
� � � � � � � � O2a2b1a1a1a3   Z25907, Z25908, Z25909
� � � � � � � � O2a2b1a1a2   FGC23469/Z25852
� � � �  � � � � O2a2b1a1a2a   F310, F666, F692
� � � �  � � � � O2a2b1a1a2a1   F402, F624
� � � � � � � �  � � O2a2b1a1a2a1a   F1531, F2781
� � � � � � � � O2a2b1a1a3   CTS7634
� � � � � � � � O2a2b1a1a3a   F317, F375, F474, F574, F869, F1401, F1707, F1870, F2000, F2188, F2494, F2631, F2845, F3212, F3431, F3455, Y16475, Y16476, Y16478, Y16479, Y16696
� � � � � � � � O2a2b1a1a3a1   F3039
� � � � � � � � O2a2b1a1a3a2   Y29861, Y30150
� � � �  � � � � O2a2b1a1a3b   CTS5488, PH3164, PH3555, Z39594
� � � � � � � � O2a2b1a1a4   Z25853
� � � � � � � � O2a2b1a1a4a   CTS5492, CTS4789, CTS5063, CTS7418, CTS8299, CTS12655
� � � �  � � � � O2a2b1a1a4a1   CTS6987, Z25902, Z25903, Z25905
� � � � � � � �  � � O2a2b1a1a4a1a   Z42620
� � � �  � � � � O2a2b1a1a4a2   F20963, Z44926, Z44927
� � � � � � � � O2a2b1a1a5   CTS10738/M1707, CTS1017/M1532, CTS7316, M1543, M1694, M1726
� � � � � � � � O2a2b1a1a5a   CTS9678, CTS8848/M1672, CTS10804/M1708, CTS11844, M1540, M1541, M1575, M1586, M1624, M1650, M1698, M1701, M1722, M1732, Z25850
� � � �  � � � � O2a2b1a1a5a1   Z39663, Z39664
� � � �  � � � � O2a2b1a1a5a2   M1513, M1557, M1593, M1602, M1604, M1608, M1617, M1629, M1637, M1683, M1688, M1719, Z44923, Z44924, Z44925
� � � �  � � � � O2a2b1a1a5b   A9457, A9458, F16367, F24592
� � � �  � � � �  � O2a2b1a1a5b1   F17158, Z44918, Z44919, Z44921, Z44922
� � � �  � � � � O2a2b1a1a6   CTS4658, CTS1642, CTS9332
� � � �  � � � � O2a2b1a1a6a   CTS5308
� � � � � � � � O2a2b1a1a6b   Z25928, Z25929, Z25930, Z25931
� � � �  � � � � O2a2b1a1a6b1   SK1730/Z25982, Z25975, Z25981, Z25983, Z25984, Z25985, Z26000
� � � � � � � �  � � O2a2b1a1a6b1a   Z26030, Z26031
� � � �  � � � � � � O2a2b1a1a6b1b   Z26010, Z26012, Z26013, Z26014
� � � � � � � � O2a2b1a1a6b2   A9462, A9461, A9463, A9464, A9465, A9466, A9467, A9468
� � � � � � � � O2a2b1a1a6b3   B456, Z39703, Z39704, Z39705, Z39706, Z39707, Z39708
� � � �  � � � � O2a2b1a1a7   YP4864, YP4865, Z44052, Z44053, Z44054, Z44055, Z44056, Z44057, Z44058, Z44059, Z44060, Z44061, Z44062, Z44063, Z44064, Z44065, Z44066, Z44067
� � � �  � � � �  O2a2b1a1a7a   Z44068, Z44069
� � � �  �  �  O2a2b1a1a7a~ F5524/SK1746, Z44070
� � � �  �  �  � O2a2b1a1a7a1~ F5525/SK1748
� � � �  � � � �  O2a2b1a1a7b   Z44071, Z44072, Z44073, Z44074, Z44075, Z44076, Z44077, Z44078, Z44079, Z44080, Z44081, Z44082, Z44083, Z44084, Z44085, Z44086, Z44087, Z44088, Z44089, Z44090
� � � �  � � � � O2a2b1a1a8   Z44091
� � � �  � � � �  O2a2b1a1a8a   Z44092, Z44093, Z44094
� � � �  � � � O2a2b1a1b   CTS4960, SK1717, SK1720, SK1721
� � � � � � O2a2b1a2   F114, CTS7655, F122, F274/Page125, F403, F430, F444, F499/Y19, F593, F701
� � � �  � � � O2a2b1a2a   F79, F314, F372, F629
� � � �  � � � � O2a2b1a2a1   F46/Y15, CTS2643, CTS11192, CTS11459, F174, F323/Y17, F353, F388, F442, F502, F507/Y18, F532, F2039
� � � � � � � � O2a2b1a2a1a   FGC16847/Z26091
� � � � � � � � O2a2b1a2a1a1   F48, CTS326, CTS816, CTS1011, CTS3871, CTS11177, F163, F275, F523, F575
� � � � � � � � � � O2a2b1a2a1a1a   F152, CTS1098, CTS8380, CTS9172, CTS9217, F279, F595, F700
� � � � � � � � � � � O2a2b1a2a1a1a1   F2505
� � � � � � � � � � O2a2b1a2a1a1b   CTS3149, CTS5323, CTS5443, CTS12551
� � � � � � � � O2a2b1a2a1a2   F242, CTS6493, CTS10286, CTS10401, CTS10888, CTS11492, F273, F563, F634, F1116
� � � � � � � � � � O2a2b1a2a1a2a   CTS4266
� � � � � � � � � � � O2a2b1a2a1a2a1   Z26108, Z26109, Z26110, Z26111, Z26112, Z26113
� � � � � � � � � � � � O2a2b1a2a1a2a1a   F2173, F1458, F3387, F3580, F3614
� � � �  � � � � O2a2b1a2a1a3   F2887, CTS3776, CTS9862
� � � � � � � � � � O2a2b1a2a1a3a   F3607
� � � � � � � � � � � O2a2b1a2a1a3a1   F3525
� � � � � � � � � � O2a2b1a2a1a3b   CTS3763, FGC16857
� � � �  � � � � � � � O2a2b1a2a1a3b1   A9472
� � � �  � � � � � � � O2a2b1a2a1a3b2   FGC16863/Y7110
� � � � � � � �  � � � � O2a2b1a2a1a3b2a   L1360, FGC16864, FGC16866, FGC16867/Y13527, FGC16869/Y13528, FGC16948
� � � �  � � � � � � � � O2a2b1a2a1a3b2a1   FGC16889, FGC16891, FGC16913, FGC16946
� � � �  � � � � � � � � O2a2b1a2a1a3b2b   SK1768/Y7112/Z26257, CTS2083, CTS2119, CTS7323, FGC23839/Y7114/Z26259, FGC23841/Y7116/Z26268, Y7111/Z26256, Z26266
� � � � � � � � � � � � O2a2b1a2a1a3b2b1   F4249, FGC1255, FGC23844, FGC23850, FGC23852, FGC23854, FGC23855, FGC23856, FGC23857, FGC23858, FGC23860, FGC23862, FGC23863, FGC23864, FGC23865, FGC23870, FGC23871, FGC23873, SK1769, SK1770, Y16776, Y16777, Y16778, Y16779
� � � �  � � � �  � � � � O2a2b1a2a1a3b2b1a   FGC23868
� � � �  � � � �  � � � � O2a2b1a2a1a3b2b2   CTS335, CTS558, CTS2080.2, CTS2882, CTS3249, CTS5450, CTS7951, CTS8942, CTS10883, CTS11454
� � � � � � � � O2a2b1a2a1b   CTS53, CTS448, CTS610, CTS4696, CTS10819, CTS11910, SK1756/Z26318
� � � � � � � � O2a2b1a2a1b1   CTS6373
� � � � � � � �  � � � O2a2b1a2a1b1a   A9473
� � � � � � � � O2a2b1a2a1c   F3386
� � � � � � � � O2a2b1a2a1d   Y29828, Y30477, Y30485, Y30487, Y30489, Y30493, Y30498, ZS11791
� � � � � � � �  � O2a2b1a2a1d1   F735, F2903, L932.2, Y29836, Y30471, Y30484, Y30499
� � � � � � � �  � � O2a2b1a2a1d1a   FGC34973, Y29829, Y29830, Y29831, Y29832, Y29833, Y29834, Y29835, Y30465, Y30466, Y30467, Y30468, Y30469, Y30470, Y30472, Y30473, Y30474, Y30475, Y30476, Y30478, Y30479, Y30480, Y30481, Y30482, Y30483, Y30486, Y30488, Y30490, Y30491, Y30492, Y30494, Y30495, Y30496, Y30497, Y30500
� � � � � � � �  � � O2a2b1a2a1d1b~   F1739
� � � � � � � O2a2b1a2b   F743, CTS1933, CTS2260, CTS8043, CTS10219, F1725, F1835, F2599, F2600, F3451  F14400/Z26072, F14403/Z26073, F14487/Z26074, F14497/Z26075, F14503/Z26076, F14559/Z26077, F14690/Z26080, F14695/Z26081, F14714/Z26083, FGC61160, FGC61167, Y31256, Z26078 
� � � � � � � � O2a2b1a2b1   CTS8481, CTS2205, FGC61159, FGC61164, FGC61165, FGC61215, Y31257
� � � �  � � � �  O2a2b1a2b1a   CTS4325, A16615, A16616, A16617, A16618, A16621, A16622, A16623, A16624, A16625, A16626, A16628, CTS496, CTS724, CTS1707, CTS2147, CTS2390, CTS4336, CTS5529, CTS7277, CTS8451, CTS9362, CTS11009
� � � �  � � � �  � O2a2b1a2b1a1   A16629, A16630
� � � �  � � � �  � O2a2b1a2b1a2   CTS682, CTS2272, CTS7184, CTS8951
� � � �  � � � � O2a2b1a2b2   F748, F823, F1448, F2093, F2095, F2212, F2236, F2530, F2795, F2974, F3279, F3383, F3440, F3505, F3579
� � � � � � � � O2a2b1a2b2a   F728, F1060, F2018, F2326
� � � � � � � O2a2b1a2c~ Page101
� � � � O2a2b2   AM01822/F3223, AM01747/F1672, AM01749/F1693, AM01770/F2029, AM01776/F2083, AM01777/F2139, AM01788/F2469, AM01800/F2683, AM01823/CTS1366/F3237, AM01828/N6, F996, F1481, F1598/S3543
� � � � O2a2b2a   AM01856/F871
� � � �  � � O2a2b2a1   N7, F2472, F4079, F4286, F4328, F4329, F4333, Z25701, Z25708, Z25713, Z25716, Z25720, Z25721, Z25728, Z25730, Z25737, Z44816
� � � �  � � � O2a2b2a1a   F4110, F4144, CTS12480, F4215, F20220, F20394, F24749, Z44734, Z44735, Z44736, Z44737, Z44738, Z44739
� � � � � � � � O2a2b2a1a1   F4068F4139, F4159, F4190, F4202, F4241, F4276, F4293, F4332, F4335
� � � � � � � � O2a2b2a1a2   SK1780, SK1781
� � � �  � � � O2a2b2a1b   F4124, SK1778/Z25746, Z25747, Z25748, Z25749, Z25750, Z25752, Z25754, Z25755, Z25756, Z25757, Z25758, Z25759, Z25760, Z25761, Z25762, Z25763, Z25764, Z25765, Z25767
� � � � � � � � O2a2b2a1b1   IMS-JST008425p6, A16609, A16610, F24792, F26088, FGC14331.2/ZS4034.2, FGC28699, SK1777, Z8263, Z40733, Z44740, Z44741, Z44742, Z44743, Z44744, Z44745, Z44746, Z44747, Z44748, Z44749, Z44750, Z44751, Z44752, Z44753, Z44754, Z44755, Z44756, Z44757, Z44758, Z44759, Z44760, Z44761, Z44762, Z44763, Z44764
� � � � � � � � O2a2b2a1b2   BY15188, F14875, F15739, F15692, F15973, F16170, F17339, F18455, F18521, F18757, F19055, F20227, F20235, F20251, F20767, F21091, F21275, F22909, F23173, F23473, F23524, F23679, F24517, F24605, F25036, F25041, F25079, F25381, F25411, F25440, F25789, F25924, F26193, F26484, F26752, Z44765, Z44766, Z44767, Z44768, Z44769, Z44770, Z44771, Z44772, Z44773, Z44774, Z44775, Z44776, Z44777, Z44778, Z44779, Z44780, Z44781, Z44782, Z44783, Z44784, Z44785, Z44786, Z44787  
� � � � � � � �  O2a2b2a1b2a  F16411, F17835, F19270, F19578, F22244, F22838, F23276, F23329, F24971, Z44792
� � � �  � � O2a2b2a2   AM01845/F706, AM01755, AM01758/F1903, AM01768, AM01780, AM01790/F2486, AM01791, AM01792, AM01799/F2654, AM01801, AM01804, AM01814/F3069, AM01842, AM01857, F1645, F16175, Z44817
� � � � � � � O2a2b2a2a   F717, F919, F976, F999, F1010, F1375, F1412, F1884, F1889, F2132, F2511, F2558. F2724, F2929, F3066, F3095, F3230, F3473, F3820, SK1782, SK1784
� � � � � � � � O2a2b2a2a1   F3612, F3819
� � � � � � � � O2a2b2a2a2   SK1783
� � � � � � � O2a2b2a2b   AM01847/B451, AM01750, AM01759, AM01760, AM01761/SK1787, AM01762, AM01763, AM01771, AM01775, AM01778, AM01781/S18967/SK1786, AM01785, AM01786, AM01787, AM01794, AM01798/SK1788, AM01802, AM01805, AM01806, AM01812, AM01813, AM01815, AM01818, AM01821, AM01824, AM01827, AM01835, AM01838, AM01839, AM01844, AM01855, AM01859, AM01860, F15980, F16102, F16868, F17803
� � � � � � � � O2a2b2a2b1   A17418, A17419, A17420, A17422, A17424, A17425, A17426, A17427, A17428, A17429, A17430, A17431, A17432, A17433, A17434, A17436, A17437, A17438, A17439, A17441, A17442, A17443, A17444, A17445, A17446, A17447, A17448, A17449, A17450, A17451, YP6194 
� � � � � � � � O2a2b2a2b2   AM01756, AM01757, AM01779, AM01803, AM01810, AM01826, AM01834, AM01861, F16663, F16771, F16783 
� � � � � � � �  O2a2b2a2b2a   B450, AM00474/SK1791, AM00475/SK1792, AM00476, F16769
� � � � � � � �  O2a2b2a2b2b   AM00472/B452, AM00463, AM00464, AM00465, AM00466, AM00467, AM00468, AM00469, AM00470, AM00471, F16088, F16616, F16997, F17764, F26682
� � � � � � � �  � O2a2b2a2b2b1   F18942, F21615, F25531
� � � � � � � �  O2a2b2a2b2c   A16427, A16428, A16429, A16430, A16431, A16432
� � � � O2a2b2b   A16433, A16434, A16435, A16437 
� � � �  O2a2b2b1   A16438, SK1773
� � � �  O2a2b2b1a   SK1775, SK1776
� � � �  O2a2b2b1a1   SK1774 
� � � �  O2a2b2b1b   A16440, A16442, A16443, A16444, A16445
� � � O2a3~ M300
� � � O2a4~ M333J
� � O2b   F742, CTS1754/F4354, CTS5152, CTS6677, F1451, F1986, F2240, F4161, F4234
� � � O2b1   F1150, F1578, F1886, F4081, F4191, F4214, F4236, PF4440
� � � � O2b1a   F837, F1770, F2244, F2247, F3338
� � � �  O2b1a1   F1025  
� � � O2b2   F1055, F1238, F2051, F2944, F4098, F4178, F4179, F4194, F4238, F4245, F4279  
� � � � O2b2a   F3021, F4050, F4061, F4323

NOTES: