Y-DNA Haplogroup E and its Subclades - 2015

The entire work is identified by the Version Number and date given on the Main Page.   Directions for citing the document are given at the bottom of the Main Page.
Version History     Last revision date for this specific page: 4 November 2015

Because of continuing research, the structure of the Y-DNA Haplogroup Tree changes and ISOGG does its best to keep the tree updated with the latest developments in the field. The viewer may observe other versions of the tree on the Web. Email Ray Banks if the differences need clarification or if you find any broken links on this page.

LINKS:  Main Page   Y-DNA Tree Trunk   SNP Index   Papers/Presentations Cited   Glossary   Listing Criteria
SNP SYMBOLS:  Not on 2014 tree  Confirmed within subclade  Provisional  Private  Investigation 

The criteria for a representative SNP printed in bold for a subclade is: traditional usage, testing one in multiple labs, and/or being found in the area of the chromosome used in recent research studies.

SNPs listed below in italics (colored black or red) are quality variants from next-generation sequencing reports consistently showing as representing that subgroup.

Paragroups, subclades ending with an asterisk (*) indicate that some individuals do not test positive for any snps downstream. Since this fact is commonly known, paragroups are being omitted to simplify the display of SNPs.

Contact Person for Haplogroup E: Aaron R. Brown

Link to Experimental Composite Y-DNA Haplogroup E Tree by Ray Banks.

E   M96/PF1823, CTS433/M5384/PF1504, CTS527/M5385/PF1505, CTS860/M5388/PF1511, CTS939/M5389, CTS1008/M5391/PF1516, CTS1074/M5393/PF1519, CTS1105/M5397, CTS1262/M5399/PF1522, CTS2401/M5448/PF1633, CTS2496/M5449/PF1636, CTS2893/M5453, CTS3199/M5455/PF1649, CTS3337/PF1653, CTS3749/M5461/PF1660, CTS3774/M5462/PF1661, CTS4373/M5466/PF1669, CTS4685/M5467/PF1671, CTS4994/M5468, CTS5316/M5470/PF1683, CTS6048/M5475/PF1697, CTS6513/M5483/PF1710, CTS6755/M5485/PF1713, CTS6818/M5487, CTS7138/M5492/PF1723, CTS7824/M5495/PF1732, CTS8053/M5497/PF1738, CTS8189/M5500, CTS8269/M5501/PF1743, CTS8631/L794/M5505/PF1751, CTS8754/M5506/PF1752, CTS9440/M5510/PF1769, CTS9663/M5514/PF1774, CTS9799/M5516/PF1777, CTS10296/M5519/PF1780, CTS10330/M5520/PF1781, CTS10344/M5521/PF1783, CTS10894/M5552, CTS11216/M5556/PF1882, CTS11504/M5559/PF1892, CTS11599/PF1893, CTS11622/PF1894, CTS11644/PF1895, CTS11681/PF1896, CTS11811/M5560/PF1899, CTS11916/M5562/PF1901, CTS12798/M5574/PF1933, FGC2079, FGC6230, L339, L504/PF1803, L507/PF1874, L511/PF1888, L537/PF1507, L614/PF1898/YSC0000032, L856/PF1553, M40/PF1445/SRY4064/SRY8299, M5382/PF1502, M5396, M5403/PF1530, M5406/PF1539, M5410/PF1545, M5416/PF1552, M5417/PF1554, M5418/PF1555, M5422/PF1561, M5424/PF1563, M5425/PF2323, M5426/PF1564, M5428, M5431/PF1576, M5434/PF1581, M5436/PF1583, M5438, M5439/PF1590, M5440/PF1591, M5452/PF1642, M5525/PF1795, M5527/PF1796, M5529/PF1798, M5531, M5533, M5539, M5540/PF1816, M5543/PF1819, M5545/PF1822, M5548/PF1834, M5549/PF1837, M5550/PF1873, M5564/PF1904, M5566/PF1906, M5567/PF1918, M5569/PF1919, M5571, M5572/PF1923, M9051, M9052, P29/PF1643, P150/PF1704, P152/PF1646, P154/PF1788, P155/PF1687, P156/PF1725, P162/PF1496, P168, P169/Page54/PF1878, P170/PF1656, P171/PF1903, P172/PF1510, P173, P174/PF1674, P175, P176/PF1678, PF1459, PF1472, PF1473, PF1476, PF1480, PF1495, PF1497, PF1501, PF1567, PF1603, PF1608, PF1618, PF1620, PF1843, PF1844, PF1864, PF1867, PF1868, PF1917, Z15560, Z15661, Z15662, Z15663, Z15664, Z15665, Z15667, Z15668, Z15669, Z15670, Z15671, Z15672, Z15673, Z15674, Z15677, Z15680, Z15681, Z15682, Z15683
E1   P147/PF1938, CTS955/M5390/PF1514, CTS5913/M5474/PF1694, CTS6355/M5479/PF1703, CTS8432/PF1745, CTS9083/M5509/PF1764, CTS9753/M5515/PF1776, M5413/PF1549, M5421/PF1559, M5534/PF1809, PF1593, Z15558, Z15559
• • E1a   M132, CTS21, CTS140, CTS245, CTS340, CTS443, CTS520, CTS548, CTS600, CTS685, CTS895, CTS1059, CTS1076, CTS1134, CTS1172, CTS1235/Z903, CTS1364, CTS1374, CTS1404, CTS1559, CTS1571, CTS1671/Z924, CTS1686/Z925, CTS1720, CTS1876, CTS1923, CTS1931, CTS1937/Z926, CTS1987, CTS2002, CTS2031, CTS2054, CTS2131/Z927, CTS2166/Z928, CTS2186, CTS2255, CTS2280, CTS2362, CTS2384/Z929, CTS2408, CTS2676, CTS2743, CTS2751, CTS2808/L633, CTS3003, CTS3045/Z930, CTS3173, CTS3309, CTS3342, CTS3459, CTS3507, CTS3581, CTS3670, CTS3678, CTS3698, CTS3714, CTS3716, CTS4034, CTS4268, CTS4453, CTS4467, CTS4477, CTS4591/Z933, CTS4946, CTS5009, CTS5033, CTS5138, CTS5194, CTS5227, CTS5283, CTS5313/Z935, CTS5393/Z936, CTS5503/Z937, CTS5589, CTS5701, CTS5734, CTS5775, CTS5793, CTS5905, CTS6063, CTS6213, CTS6230, CTS6245, CTS6288, CTS6370, CTS6410, CTS6476, CTS6486, CTS6516, CTS6608/Z939, CTS6614/Z940, CTS7156, CTS7190, CTS7296, CTS7584, CTS7651, CTS7654, CTS7830, CTS8185, CTS8249, CTS8383, CTS8433, CTS8465, CTS8482, CTS8538, CTS8632, CTS8663, CTS8720, CTS8835, CTS8867, CTS8953, CTS9003, CTS9074, CTS9110, CTS9114/Z942, CTS9155, CTS9157, CTS9198, CTS9439, CTS9455, CTS9505/Z943, CTS9520, CTS9673, CTS9714, CTS9927, CTS10067, CTS10147, CTS10182, CTS10475, CTS10497, CTS10567, CTS10644/Z957, CTS10752, CTS10964, CTS11187, CTS11248, CTS11276/Z960, CTS11300/Z961, CTS11329, CTS11444, CTS11591, CTS11860, CTS12046/Z962, CTS12271, CTS12333, CTS12743, L481, M9060, M9061/S12552, M9062/S12553, M10372, M33, Y1193.2, Z875, Z883, Z898, Z899, Z900, Z901, Z902, Z904, Z907, Z910, Z911, Z912, Z913, Z916, Z917, Z920, Z921, Z922, Z931, Z938, Z944, Z945, Z946, Z948, Z949, Z950, Z951, Z953, Z954, Z963, Z964, Z966, Z15412, Z15413, Z15414, Z15415, Z15416, Z15417, Z15418, Z15419, Z15420, Z15421, Z15422, Z15423, Z15424, Z15425, Z15426, Z15427, Z15428, Z15429, Z15430, Z15431, Z15432, Z15433, Z15434, Z15435, Z15436, Z15437, Z15438, Z15439, Z15440, Z15441, Z15442, Z15443, Z15444, Z15445, Z15446, Z15447, Z15448, Z15449, Z15450, Z15451, Z15452, Z15453, Z15454, Z15455, Z15456, Z15457, Z15458, Z15459, Z15460, Z15461, Z15462, Z15463, Z15464, Z15465/Z17396, Z15466, Z15467, Z15468, Z15469, Z15470, Z15471, Z15472, Z15473, Z15474, Z15475, Z15476, Z15477, Z15478, Z15479, Z15480, Z15481, Z15482, Z15483, Z15484, Z15485, Z15486, Z15487, Z15488, Z15489, Z15490, Z15491, Z15492, Z15493, Z15494, Z15495, Z15496, Z15497, Z15498, Z15499, Z15500/Z17398, Z15501, Z15502, Z15503, Z15504, Z15505, Z15506, Z15507, Z15508, Z15509, Z15510, Z15511, Z15512, Z15513, Z15514, Z15515, Z15516, Z15517/Z17400, Z15518, Z15519, Z15520, Z15521, Z15522, Z15523, Z15524, Z15525, Z15526, Z15527, Z15528, Z15529, Z15530, Z15531, Z15532, Z15533, Z15534, Z15535, Z15536, Z15537, Z15538, Z15539, Z15540, Z15541, Z15542, Z15543, Z15544, Z15545, Z15546, Z15547, Z15548, Z15549, Z15550, Z15551, Z15552, Z15553, Z15554, Z15555, Z15556, Z15557, Z17397, Z17399, Z17401, Z17402, Z17403
• • • E1a1   M44, CTS1276/M11794/PF2422, BY172.1/SK563.1/Z17470.1, L632, L634, L1238, L1239, L1240, SK562/Z17420, SK564/Z17450, SK565/Z17480, SK566/Z17422, SK567/Z17417, SK568/Z17498, Z17405, Z17406, Z17407, Z17408, Z17409, Z17410, Z17411, Z17412, Z17413, Z17414, Z17415, Z17416, Z17418, Z17419, Z17421, Z17423, Z17424, Z17425, Z17426, Z17427, Z17428, Z17429, Z17430, Z17431, Z17432, Z17433, Z17434, Z17435, Z17436, Z17437, Z17438, Z17439, Z17440, Z17441, Z17442, Z17443, Z17444, Z17445, Z17446, Z17447, Z17448, Z17449, Z17451, Z17452, Z17453, Z17454, Z17455, Z17456, Z17457, Z17458, Z17459, Z17460, Z17461, Z17462, Z17463, Z17464, Z17465, Z17466, Z17468, Z17469, Z17471, Z17472, Z17473, Z17474, Z17475, Z17476, Z17477, Z17478, Z17479, Z17481, Z17482, Z17483, Z17484, Z17485, Z17486, Z17487, Z17488, Z17489, Z17490, Z17491, Z17492, Z17493, Z17495, Z17496, Z17497, Z17499, Z17500, Z17501, Z17502, Z17503, Z17504, Z17505, Z17506, Z17507
• • • • E1a1a   Z17699
• • • •  E1a1a1   Z17697, Z17698, Z17700
• • • •  E1a1a1a   Z17696
• • • •  E1a1a2   A7710
• • • • E1a1b   Z17467
• • • •  E1a1b1   Z31503
• • • E1a2   CTS10713/Z958, CTS230, CTS248, CTS678, CTS986, CTS1103, CTS1306, CTS1319, CTS1556, CTS2223, CTS2307, CTS2327, CTS2454, CTS2832, CTS3025, CTS3208, CTS3256, CTS3355, CTS3515, CTS3538, CTS3914, CTS4174/Z932, CTS4276, CTS4316, CTS4337, CTS4692, CTS5155/Z934, CTS5422, CTS5458, CTS6239, CTS6384, CTS6817, CTS6891, CTS7198, CTS7453/Z941, CTS7499, CTS7851, CTS7978, CTS8133, CTS8817, CTS9096, CTS9212, CTS9548, CTS10225, CTS10801, CTS11085/Z959, CTS11320, CTS11390, CTS12022, CTS12067, CTS12129, PF3085, Z868, Z869, Z877, Z897, Z905, Z908, Z914, Z915, Z952, Z955, Z956, Z965, Z967, Z15106, Z15107, Z15108, Z15109, Z15110, Z15111, Z15112, Z15113, Z15114, Z15115, Z15116, Z15117, Z15118, Z15119, Z15120, Z15121, Z15122, Z15123, Z15124, Z15125, Z15126, Z15127, Z15128, Z15129, Z15130, Z15131, Z15132, Z15133, Z15134, Z15135, Z15136, Z15137, Z15138, Z15139, Z15140, Z15141, Z15142, Z15143, Z15144, Z15145, Z15146, Z15147, Z15148, Z15149, Z15150, Z15151, Z15152, Z15153, Z15154, Z15155, Z15156, Z15157, Z15158, Z15159, Z15160, Z15161, Z15162, Z15163, Z15164, Z15165, Z15166, Z15167, Z15169
• • • • E1a2a   CTS246, CTS2632, CTS4519, CTS9089, Y967, Z906, Z918, Z15171
• • • •  E1a2a1   CTS10935, Z15271
• • • •  E1a2a1a   CTS3380, CTS2648, CTS4187, Z885
• • • •  • • E1a2a1a1   P110, CTS1753, CTS1927/Z888, CTS2273, CTS2518/Z889, CTS3519, CTS3881, CTS5110/Z1890, CTS5365, CTS7182, CTS8725, CTS11593, CTS11290/Z894, CTS11578/Z895, Z884, Z886, Z887, Z892, Z15395, Z15396
• • • •  • • • E1a2a1a1a   CTS1792, CTS6482, CTS7040, CTS7797, CTS10874, Z15397, Z15398, Z15399, Z15400, Z15401, Z15402, Z15403, Z15404, Z15405
• • • •  • • E1a2a1a2   Z5993, Z15406, Z15407, Z15408, Z15409, Z15410, Z15411
• • • •  E1a2a1b   L133.1/Page74.1, CTS1285, CTS5145/Z880, CTS5607, CTS6573, CTS7767.2, CTS8762/Z881, CTS11769, Z878, Z879, Z15272, Z15273, Z15274, Z15275, Z15276, Z15277, Z15278
• • • •  • • E1a2a1b1   CTS736, A173.1/Z15291.1, CTS602, CTS3351, CTS3615, CTS3840, CTS7084, CTS7140, CTS7912, CTS9489/M1116, CTS9792, CTS10712, CTS11737, Z12510.2/Z15303.2, Z15279, Z15280, Z15281, Z15282, Z15283, Z15284, Z15285, Z15286, Z15287, Z15288, Z15289, Z15290, Z15292, Z15293, Z15294, Z15295, Z15296, Z15297, Z15298, Z15299, Z15300, Z15301, Z15302, Z15304, Z15305, Z15306, Z15307, Z15308, Z15309, Z15310, Z15311, Z15312, Z15313, Z15314, Z15315, Z15316, Z15317, Z15318, Z15319, Z15320, Z15321, Z15322, Z15323, Z15324, Z15325, Z15326
• • • •  • • • E1a2a1b1a   CTS3764.1, Z5992, Z15327, Z15328, Z15329, Z15330, Z15331, Z15332, Z15333, Z15334, Z15335, Z15336, Z15337, Z15338, Z15339, Z15340, Z15341, Z15342, Z15343, Z15344, Z15345, Z15346, Z15347, Z15348, Z15349, Z15350, Z15351, Z15352, Z15353, Z15354, Z15355, Z15356, Z15357, Z15358, Z15359, Z15360, Z15361, Z15362, Z15363, Z15364, Z15365, Z15366, Z15367, Z15368, Z15369, Z15370, Z15371, Z15372, Z15373, Z15374, Z15375, Z15376, Z15377, Z15378, Z15379, Z15380, Z15381, Z15382, Z15383, Z15384, Z15385, Z15386, Z15387, Z15388, Z15389, Z15390, Z15391, Z15392, Z15393, Z15394
• • • •  E1a2a2   Z5991, Z15172, Z15173, Z15174, Z15175, Z15176, Z15177, Z15178, Z15179, Z15180, Z15181, Z15182, Z15183, Z15184, Z15185, Z15186, Z15187, Z15188, Z15189, Z15190, Z15191, Z15192, Z15193, Z15194, Z15195, Z15196, Z15197, Z15198, Z15199, Z15200, Z15201, Z15202, Z15203, Z15204, Z15205, Z15206, Z15207, Z15208, Z15209, Z15210, Z15211, Z15212, Z15213, Z15214, Z15215, Z15216, Z15217, Z15218, Z15219, Z15220, Z15221, Z15222, Z15223, Z15224, Z15225, Z15226, Z15227, Z15228, Z15229, Z15230, Z15231, Z15232, Z15233, Z15234, Z15235, Z15236, Z15237, Z15238, Z15239, Z15240, Z15241, Z15242, Z15243, Z15244, Z15245, Z15246, Z15247, Z15248, Z15249, Z15250, Z15251, Z15252, Z15253, Z15254, Z15255, Z15256, Z15257, Z15258, Z15259, Z15260, Z15261, Z15262, Z15263, Z15264, Z15265, Z15266, Z15267, Z15268, Z15269, Z15270
• • • • E1a2b   Z5985.1, Z15084, Z15085, Z15086, Z15087, Z15088, Z15089, Z15090, Z15091, Z15092, Z15093, Z15094, Z15095, Z15096, Z15097, Z15098, Z15099, Z15100, Z15101, Z15102, Z15103, Z15104, Z15105
• • • •  E1a2b1   L94, Z15068, Z15069, Z15070, Z15071, Z15072, Z15073, Z15074, Z15075, Z15076, Z15077, Z15078, Z15079, Z15080, Z15081, Z15082, Z15083
• • • •  E1a2b1a   Z5986, Z15062, Z15063, Z15064, Z15065, Z15066, Z15067
• • • •  • • E1a2b1a1   Z5988
• • • •  • • • E1a2b1a1a   Z5989, Z14979
• • • •  • • • • E1a2b1a1a1   Z14889, Z14890, Z14891, Z14892, Z14893, Z14894, Z14895, Z14896, Z14897, Z14898, Z14899, Z14900, Z14901, Z14902, Z14903, Z14904, Z14905, Z14906, Z14907, Z14908, Z14909, Z14910, Z14911, Z14912, Z14913, Z14914, Z14915, Z14916, Z14917, Z14918, Z14919, Z14920, Z14921, Z14922, Z14923, Z14924, Z14925, Z14926, Z14927, Z14928, Z14929, Z14930, Z14931, Z14932, Z14933, Z14934, Z14935, Z14936, Z14937, Z14938, Z14939, Z14940, Z14941, Z14942, Z14943, Z14944, Z14945, Z14946, Z14947, Z14948, Z14949, Z14950, Z14951, Z14952, Z14953, Z14954, Z14955, Z14956, Z14957, Z14958, Z14959, Z14960, Z14961, Z14962, Z14963, Z14964, Z14965, Z14966, Z14967, Z14968, Z14969, Z14970, Z14971, Z14972, Z14973, Z14974, Z14975, Z14976, Z14977, Z14978
• • • •  • • E1a2b1a2   M9119, Z5987, Z14980, Z14981, Z14982, Z14983, Z14984, Z14985, Z14986, Z14987, Z14988, Z14989, Z14990, Z14991, Z14992, Z14993, Z14994, Z14995, Z14996, Z14997, Z14998, Z14999, Z15000, Z15002, Z15003, Z15004, Z15005, Z15006, Z15007, Z15008, Z15009, Z15010, Z15011, Z15012, Z15013, Z15014, Z15015, Z15016, Z15017, Z15018, Z15019, Z15020, Z15021, Z15022, Z15023, Z15024, Z15025, Z15026, Z15027, Z15028, Z15029, Z15030, Z15031, Z15032, Z15033, Z15034, Z15035, Z15036, Z15037, Z15038, Z15039, Z15040, Z15041, Z15042, Z15043, Z15044, Z15045, Z15046, Z15047, Z15048, Z15049, Z15050, Z15051, Z15052, Z15053, Z15054, Z15055.1, Z15056, Z15057, Z15058, Z15059, Z15060, Z15061, Z15170
• • E1b   P177/PF1939
• • • E1b1   DYS391p, P2/PF1940/PN2, P178, P179, P180/PF1941, P181
• • • • E1b1a   L222.1/S350.1, TSC0077541/V38, V100
• • • •  E1b1a1   CTS1445/Z1107, CTS3105, CTS3989, CTS4245/Z1120, CTS4350, CTS4415, CTS5042/Z1122, CTS5539, CTS6319/Z1124, CTS6474, CTS7095, CTS7641, CTS8068, CTS8123, CTS8443/TSC0769719, CTS8936, CTS9188, CTS9338, CTS10066, CTS10638, CTS10806, CTS11461/KHS1479188, CTS11579, CTS11729, DYS271/M2/SY81, L488, L491, L494, L608, L610, M291, P1/PN1, P189.1, P293, Page106, CTS5964/TSC0424987, V95, Z1101, Z1116, Z1123, Z1125, Z1127, Z1130, Z1133, Z1135
• • • •  E1b1a1a   P46
• • • •  • • E1b1a1a1   CTS1847, CTS3576, CTS10659, CTS10914, CTS11732/KHS1381212, CTS12659, L86.1, M180/P88, P182, Page66, V43, Z1111, Z1132
• • • •  • • • E1b1a1a1a   M58, Page27
• • • •  • • • E1b1a1a1b   M10
• • • •  • • • E1b1a1a1c    L485
• • • •  • • • • E1b1a1a1c1   L514
• • • •  • • • •  E1b1a1a1c1a   M191/P86, U186, P253/U247, TSC1640819/Z1712.1
• • • •  • • • •  E1b1a1a1c1a1   P252/U174
• • • •  • • • •  • • E1b1a1a1c1a1a   P9.2
• • • •  • • • •  • • E1b1a1a1c1a1b   P115
• • • •  • • • •  • • E1b1a1a1c1a1c   CTS8030/Z1704
• • • •  • • • •  • • • E1b1a1a1c1a1c1   CTS9106/M4451
• • • •  • • • •  • • • • E1b1a1a1c1a1c1a   P116
• • • •  • • • •  • • • •  E1b1a1a1c1a1c1a1   P113
• • • •  • • • •  • • E1b1a1a1c1a1d   CTS1313/PF7223
• • • •  • • • •  E1b1a1a1c1b   L515, L516, L517, M263.2, Z1893
• • • •  • • • •  E1b1a1a1c1b1   CTS3393/Z1894
• • • •  • • • • E1b1a1a1c2   CTS9883, CTS1546, CTS5961, CTS10560, CTS10932, CTS10996, Z15933, Z15934, Z15935, Z15936, Z15937, Z15938, Z15939, Z15940, Z15941, Z15942, Z15943, Z15944, Z15945, Z15946, Z15947, Z15948, Z15949, Z15950, Z15951, Z15952, Z15953, Z15955, Z15956, Z15958, Z15959, Z15960, Z15961, Z15962, Z15963, Z15964
• • • •  • • • •  E1b1a1a1c2a   Z6003, Z15965, Z15966, Z15967, Z15968, Z15969, Z15970, Z15971, Z15972, Z15973, Z15974, Z15975, Z15976, Z15977, Z15978, Z15979, Z15980, Z15981, Z15982, Z15983, Z15984, Z15985, Z15986
• • • •  • • • •  E1b1a1a1c2a1   Z6004, Z15988, Z15989, Z15990, Z15991, Z15992, Z15993, Z15994, Z15995, Z15996
• • • •  • • • •  E1b1a1a1c2b   Z6005, Z15997, Z15998, Z15999, Z16000, Z16001, Z16002, Z16003, Z16004, Z16005, Z16006, Z16007, Z16008, Z16009, Z16010, Z16011, Z16012
• • • •  • • • •  E1b1a1a1c2b1   Z6006, Z16013, Z16014, Z16015, Z16016, Z16017, Z16018, Z16019, Z16020, Z16021, Z16022, Z16023, Z16024, Z16025, Z16026, Z16027, Z16028, Z16029.1, Z16030, Z16031, Z16032, Z16033, Z16034, Z16035, Z16036, Z16037, Z16038, Z16039, Z16040, Z16041
• • • •  • • • •  E1b1a1a1c2b2   Z6007, Z16042, Z16043, Z16044, Z16045
• • • •  • • • •  • • E1b1a1a1c2b2a   Z16052, Z16046, Z16047, Z16048, Z16049, Z16050, Z16051, Z16053, Z16054, Z16055
• • • •  • • • •  E1b1a1a1c2c   CTS3274, Z16056
• • • •  • • • •  E1b1a1a1c2c1   CTS12004, CTS4907, CTS6973, CTS10951, CTS10959, CTS11236, Z16057, Z16058, Z16059, Z16061, Z16062, Z16063, Z16064
• • • •  • • • •  • • E1b1a1a1c2c1a   CTS3725, CTS308, CTS9325, CTS9930, Z16065, Z16066, Z16067, Z16068, Z16069, Z16070, Z16071, Z16072, Z16073, Z16074, Z16075, Z16076, Z16077, Z16078, Z16079, Z16080, Z16081, Z16082, Z16083, Z16084, Z16085, Z16086
• • • •  • • • •  • • E1b1a1a1c2c1b   Z6009, Z16087, Z16088, Z16089, Z16090, Z16091, Z16092, Z16093, Z16094, Z16095, Z16096, Z16097, Z16098, Z16099, Z16100, Z16101, Z16102, Z16990, Z16991, Z16992, Z16993, Z16994, Z16995, Z16996, Z16997, Z16998, Z16999, Z17000, Z17001, Z17002, Z17003, Z17004, Z17005, Z17006, Z17007, Z17008, Z17009, Z17010
• • • •  • • • •  E1b1a1a1c2c2   Z6010/Z17024, Z17011, Z17012, Z17013, Z17014, Z17015, Z17016, Z17017, Z17018, Z17019, Z17020, Z17021, Z17022, Z17023, Z17025, Z17026, Z17027
• • • •  • • • •  • • E1b1a1a1c2c2a   F2481, Z17028, Z17029, Z17030, Z17031, Z17032, Z17033, Z17034, Z17035, Z17036, Z17037, Z17038, Z17039, Z17040, Z17041, Z17042, Z17043, Z17044, Z17045, Z17046, Z17047, Z17048
• • • •  • • • •  E1b1a1a1c2c3   Z6012
• • • •  • • • •  • • E1b1a1a1c2c3a   Z6013
• • • •  • • • •  • • • E1b1a1a1c2c3a1   Z6014, Z17049, Z17050, Z17051, Z17052, Z17053, Z17054, Z17055, Z17056, Z17057, Z17058, Z17059, Z17060, Z17061, Z17062, Z17063, Z17064, Z17065, Z17066, Z17067, Z17068, Z17069, Z17070, Z17071, Z17072, Z17073
• • • •  • • • •  • • • E1b1a1a1c2c3a2   Z6015, M7820, Z16103, Z16104, Z16105, Z16106, Z16107, Z16108, Z16109, Z16110, Z16111, Z16112, Z17074, Z17075, Z15796.2/Z17076.2, Z17077
• • • •  • • • •  • • • • E1b1a1a1c2c3a2a   Z6016, Z16113, Z16114, Z16115, Z16116, Z16117, Z16118, Z16119, Z16120, Z16121, Z16122, Z16123, Z16124, Z16125
• • • •  • • • •  • • E1b1a1a1c2c3b   Z6017
• • • •  • • • •  • • • E1b1a1a1c2c3b1   Z6018, Z16126, Z16127, Z16128, Z16129, Z16130, Z16131, Z16132, Z16133, Z16134, Z16135, Z16136, Z16137, Z16138, Z16139, Z16140, Z16141, Z16142, Z16143, Z16144, Z16145, Z16146, Z16147, Z16148, Z16149, Z16150, Z16151, Z16152, Z16153, Z16154, Z16155, Z16156, Z16157, Z16158, Z16159, Z16160, Z16161
• • • •  • • • •  • • E1b1a1a1c2c3c   Z6019, Z16162, Z16163, Z16164, Z16165, Z16166, Z16167, Z16168, Z16169, Z16170, Z16171, Z16172, Z16173, Z16174, Z16175, Z16176, Z16177, Z16178, Z16179, Z16180, Z16181, Z16182, Z16183, Z16184, Z16185, Z16186, Z16187, Z16188, Z16189, Z16190, Z16191, Z16192, Z16193, Z16194, Z16195, Z16196, Z16197, Z16198, Z16199, Z16200, Z16201, Z16202, Z16203, Z16204, Z16205, Z16206, Z16207, Z16208, Z16209, Z16210, Z16211, Z16212, Z16213
• • • •  • • • E1b1a1a1d   U175
• • • •  • • • • E1b1a1a1d1   M4254, P277, P278.1, U209
• • • •  • • • •  E1b1a1a1d1a   U290
• • • •  • • • •  E1b1a1a1d1a1   U181
• • • •  • • • •  • • E1b1a1a1d1a1a   L97
• • • •  • • • •  E1b1a1a1d1a2   CTS99/Z1725
• • • •  • • • •  E1b1a1a1d1b   P59
• • • •  • • • •  E1b1a1a1d1c   M154
• • • •  • • • •  E1b1a1a1d1d   V39
• • • •  E1b1a1b   Z5994
• • • •  • • E1b1a1b1   Z22132
• • • •  • • • E1b1a1b1a   Z5999
• • • •  • • • • E1b1a1b1a1   Z6000
• • • •  • • • E1b1a1b1b   Z5996
• • • •  • • • E1b1a1b1c   Z5998
• • • •  E1b1a2   M329
• • • • E1b1b   M215/Page40/PF1942, CTS225/M5017/PF1450, CTS2591/M5136/PF1637, CTS3988/M5161/PF1664, CTS5530/M5183/PF1689, CTS5792/M5190/PF1692, CTS6953/M5221, CTS7677/L930/M5234/PF1729, CTS8033/M5038/PF1737, CTS8479.1/M5251.1/PF1746.1, CTS8586/M5255/PF1749, CTS9017/M5261, CTS9049/M5039/PF1763, CTS10513/M5282/PF1789, CTS10637/M5327, CTS10679/M5048/PF1875, CTS11223/L676/M5049/PF1883, L336/PF1825, M243/PF1943, M5082/PF1537, M5083/PF1538, M5084/PF1540, M5101/PF1560, M5112/PF1569, M5113/PF1570, M5299/PF1807, M5304/PF1818, M5305/PF1820, M5314/PF1826, M5316/PF1831, M5318/PF1832, M5372/PF1929, PF1484, PF1487, PF1498, PF1571, PF1572, PF1574
• • • •  E1b1b1   M35.1, CTS890/M5059/PF1513, CTS1389/M5069/PF1525, CTS2216/M5026/PF1629, CTS2474/M5134/PF1635, CTS2620/M5137/PF1638, CTS3512/M5150/PF1655, CTS3637/M5156/PF1659, CTS4220/M5166, CTS4856/M5174/PF1677, CTS6298/M5200/PF1701, CTS6809/M5213/PF1716, CTS6834/L931/PF1718, CTS7154/M5226/PF1724, CTS7980/M5241/PF1735, CTS8131/M5247/PF1740, CTS8945/M5258/PF1759, CTS9324/M5266/PF1768, CTS9956/PF1778, CTS10184/M5274/PF1779, L117/Page15/PF1657, L538/M5100/PF1556, L545/M5178/PF1679, L796/M5294/PF1801, M5021/PF1532, M5041/PF1812, M5047/PF1830, M5074/PF1531, M5078/PF1534, M5085/PF1542, M5088/PF1543, M5108/PF1566, M5287/PF1793/S7492, M5301/PF1813, M5321/PF1835, M5322/PF1836, M5325/PF1871, M5360/PF1909, M5362/PF1913, PF1454, PF1457, PF1466, PF1471, PF1492, PF1499, PF1575, PF1598, PF1619, PF1755
• • • •  E1b1b1a   V68.1, CTS2270/PF2127, CTS2661/PF2130, CTS4208/PF2136/Z1209, CTS7924/PF2154, CTS8899, CTS10323/PF2167, L539/PF2203, L546/PF2153, PF1956, PF2108, PF2114/S7500, PF2115, PF2173/Z1211, PF2175/S7478, PF2178, PF2182, PF2185/Z1214, PF2188
• • • •  • • E1b1b1a1   L18, M78/PF2186
• • • •  • • • E1b1b1a1a   Z1902
• • • •  • • • • E1b1b1a1a1   V12/Z1216, CTS154, CTS1810, CTS1833, CTS2163, CTS2402, CTS2851, CTS2967, CTS3073/Z1219, CTS3578/Z1220, CTS4185, CTS5254/Z1223, CTS5412/Z1224, CTS5646, CTS5669, CTS5752, CTS6211, CTS7017, CTS7136, CTS9007, CTS12019, CTS12269, Z1218, Z1225, Z1226
• • • •  • • • •  E1b1b1a1a1a   M224
• • • •  • • • •  E1b1b1a1a1b   V32, CTS1257/Z810, CTS3301/Z815, CTS7146/Z820, CTS10654/Z823, Z811, Z812, Z813, Z814, Z816, Z817, Z818, Z819, Z821, Z822, Z824
• • • •  • • • •  E1b1b1a1a1b1   CTS1210/Z809, Z808
• • • •  • • • •  E1b1b1a1a1c   CTS693, Z807
• • • •  • • • • E1b1b1a1a2   PF2272/V65, CTS1414/M5072/PF2317, CTS5168/PF2264, CTS6672/Z1235, CTS8686/Z1238, CTS9060/Z1239, CTS9879/Z1240, CTS10414, CTS11387/PF2295, CTS11582/Z1243, L66, L67/PF2267, PF2159, PF2253, PF2254, PF2255, PF2257, PF2258, PF2259, PF2261, PF2262, PF2263, PF2274, PF2276/Z1237, PF2286, PF2290, PF2293, PF2297
• • • •  • • • •  E1b1b1a1a2a   PF2187, PF2265, PF2284, PF2287, PF2288, PF2291/Z1241, PF2294
• • • •  • • • E1b1b1a1b   CTS4231/PF2137/Z1919, CTS4235/PF2228/Z1920
• • • •  • • • • E1b1b1a1b1   L618, CTS1773, CTS1975/PF2126, CTS3287/PF2225, CTS5291/PF2230/Z1052, CTS5527/PF2232, CTS7273/PF2239, CTS10912/PF2249, CTS11953/Z1061, PF2121, PF2215, PF2219/Z1047, PF2246
• • • •  • • • •  E1b1b1a1b1a   L142.1, L542/PF2220, Page102, PF2211/V13, PF2210/V36
• • • •  • • • •  E1b1b1a1b1a1   L17
• • • •  • • • •  E1b1b1a1b1a2   DYS643-null/L143
• • • •  • • • •  E1b1b1a1b1a3   M35.2
• • • •  • • • •  E1b1b1a1b1a4   L241
• • • •  • • • •  E1b1b1a1b1a5   L250, L251, L252
• • • •  • • • •  E1b1b1a1b1a6   L540
• • • •  • • • • E1b1b1a1b2   CTS9547/L677, CTS2548, CTS2817, CTS5479, CTS6434, CTS8892, V22
• • • •  • • • •  E1b1b1a1b2a   M148
• • • •  • • • •  E1b1b1a1b2b   L674, L199.2/L443.2/L675.2/PF5371.2
• • • •  • • • E1b1b1a1c   M521
• • • •  E1b1b1b   CTS1243/Z827
• • • •  • • E1b1b1b1   L19/PF2345/V257, CTS1042/M5061/Z1160, CTS1303/M5066/PF2314, CTS1412/M5070/PF2315, CTS1483/PF2318, CTS2885/M5140/PF2346, CTS3305/M5145/PF2347, CTS5558/Z1183, CTS5604/M5186/PF2362, CTS6444/M5205/PF2365, CTS6754/M5209/PF2368/Z1187, CTS6911/M5220/PF2372, CTS6956/M5222, CTS7795/M5236/PF2378, CTS7796/Z1193, CTS8649/M5256/PF2381, CTS11687/PF2405, L335/PF2300, M310/PF2402, M5050/PF2410, M5042/PF2395, M5130/PF2343, M5201/PF2364, M5227/PF2375, M5272/PF2387, M5298/PF2394, M5310/PF2396, M5317/PF2398, M5363/PF2411, M5366/PF2413, PF2307, PF2319/Z1162, PF2321/Z1164, PF2322/Z1166, PF2324, PF2331, PF2333/Z1170, PF2355, PF2382/Z1196, PF2383/Z1197
• • • •  • • • E1b1b1b1a   M81/PF2554
• • • •  • • • • E1b1b1b1a1   M183/Page33, CTS87/M5054/PF2472, CTS1294/Z1161, CTS1819/M5121/PF2504/Z1172, CTS3350/M5146/PF2510, CTS3415/M5147/PF2348, CTS3870/PF2513/Z1177, CTS4236/PF2514/Z1179, CTS5819/M5191/PF2520/Z1184, CTS5883/M5193/PF2363, CTS7132/M5225/PF2531, CTS8527/M5252/PF2538/Z1195, CTS10196/M5275/PF2544, CTS10321/M5040/PF2570, CTS11147/M5329/PF2560, M5008/PF2498, M5075/PF2485, M5080/PF2486, M5093/PF2489/Z1165, M5095/PF2490, M5097/PF2491, M5099/PF2493, M5102/PF2494, M5106/PF2497, M5109/PF2499/Z1169, M5110/PF2500, M5114/PF2501, M5172/PF2515, M5281/PF2545, M5284/PF2546, M5286/PF2547, M5291/PF2548, M5307/PF2551, M5308/PF2552, M5311/PF2553, M5324/PF2401/Z1201, PF1453, PF1585, PF2301, PF2305, PF2306, PF2310/Z1158, PF2311, PF2326, PF2475, PF2476, PF2477/Z1156, PF2478, PF2479/Z1157, PF2506, PF2549, PF2557, Z826
• • • •  • • • •  E1b1b1b1a1a   M165
• • • •  • • E1b1b1b2   PF1961/Z830, CTS689/Z829, CTS3457, CTS5487/PF1979/Z832, CTS5489/Z833, CTS6504/PF1983/Z834, CTS6729/Z835, CTS8182, CTS8790/PF1989, CTS10298, CTS10531/PF1995/Z837, CTS11781/PF2006, PF1945/Z828, PF1957, PF1964, PF1965/Z1076, PF1969/Z831, PF1997, PF2002, PF2009, Z836
• • • •  • • • E1b1b1b2a   M123/PF2023, CTS677/PF2015, CTS1229, CTS1652/Z1152, CTS6404, CTS9588, CTS10656, CTS11004, L798, L799, L857/PF2017/Z1147, PF2021/Z1154
• • • •  • • • • E1b1b1b2a1   M34/PF2022, CTS1264/Z843, CTS1510/Z844, CTS10814/Z859, CTS10815/Z860, CTS10896/Z861, CTS12700/Z863, L787, L797/PF2016/Z1146, PF1999/Z1155, PF2018/Z1148, PF2020/Z1153, Z839, Z840, Z841, Z845, Z846, Z847, Z848, Z855, Z856, Z857, Z858, Z862
• • • •  • • • •  E1b1b1b2a1a   M84, CTS1096, L795/CTS4483, L29/Page47, PF6750
• • • •  • • • •  E1b1b1b2a1a1   M136
• • • •  • • • •  E1b1b1b2a1b   M290
• • • •  • • • •  E1b1b1b2a1c   V23  (V23 relationship to M84 and M290 uncertain)
• • • •  • • • •  E1b1b1b2a1d   L791, L792, CTS2074, CTS5314, CTS6147, CTS6837/Z851, CTS302/Z838, CTS8037/Z852, CTS8723/Z853
• • • •  • • • E1b1b1b2b   M293, CTS1177, CTS2297, CTS10059/Z1274, CTS10880, CTS10942, CTS11025, CTS11051, CTS11574/Z1283, Z864, Z865
• • • •  • • • • E1b1b1b2b1   P72
• • • •  • • • E1b1b1b2c    V42
• • • •  E1b1b1c   V6
• • • •  E1b1b1d   V92
• • • •  E1b1b2   M281, V16
• • • E1b2   P75
E2   M75, CTS20, CTS72, CTS249, CTS309, CTS341, CTS396, CTS471, CTS472/Z996, CTS714, CTS844/Z975, CTS918, CTS1082, CTS1173, CTS1385/Z997, CTS1430, CTS1633, CTS1986, CTS2059, CTS2113, CTS2574, CTS2590, CTS2838, CTS2912, CTS2954, CTS3009, CTS3074, CTS3373/Z1009, CTS3464/Z1010, CTS3558, CTS3643, CTS3845, CTS4007, CTS4076/Z1011, CTS4298, CTS4418, CTS4751/Z1012, CTS4847, CTS5025, CTS5133, CTS5148, CTS5436/Z1015, CTS5989, CTS6419, CTS6565, CTS6588, CTS6651, CTS6731, CTS6971, CTS7088, CTS7458, CTS7888, CTS8331, CTS8550, CTS8752, CTS8958, CTS8990/Z1020, CTS9276, CTS9424, CTS9480, CTS9536, CTS9553, CTS9579, CTS9637, CTS9818, CTS10020, CTS10216, CTS10361/Z1024, CTS10370, CTS10460, CTS10565, CTS10606, CTS10614, CTS10837, CTS11304/M10999, CTS11397, CTS11446, CTS11447/Z1034, CTS11478, CTS11653, CTS11945, CTS11960, CTS12049, CTS12124, CTS12172, CTS12242, CTS12588, CTS12701, CTS12745, CTS12780, CTS12905, CTS13023, P68, SK960/Z15727, Z988, Z995, Z999, Z1003, Z1005, Z1027, Z1028, Z15684, Z15685, Z15686, Z15687, Z15688, Z15689, Z15690, Z15691, Z15692, Z15693, Z15694, Z15695, Z15696, Z15697, Z15698, Z15699, Z15700, Z15701, Z15702, Z15703, Z15704, Z15705, Z15706, Z15707, Z15708, Z15709, Z15710, Z15711, Z15713, Z15714, Z15715, Z15716, Z15717, Z15718, Z15719, Z15720, Z15721, Z15722, Z15723, Z15724, Z15725, Z15726, Z15728, Z15729, Z15730, Z15731, Z15732, Z15733, Z15734, Z15735, Z15736, Z15737, Z15738, Z15739, Z15740, Z15741, Z15742, Z15743, Z15744, Z15747, Z15748, Z15750, Z15751, Z15752, Z15753, Z15754, Z15755, Z15756, Z15757, Z15758, Z15759, Z15760, Z15761, Z15764, Z15765, Z15766, Z15767
• • E2a   M41/P210
• • E2b   M98, CTS16, CTS98, CTS132, CTS375, CTS515, CTS1191, CTS1248, CTS1398, CTS1859, CTS1909, CTS1955, CTS2145, CTS2239, CTS2300/Z1007, CTS2564, CTS2614, CTS2696, CTS2984, CTS3092, CTS3291/Z1008, CTS3329, CTS3445, CTS3666, CTS3677, CTS3712, CTS3713, CTS3721, CTS3782, CTS3905, CTS4310, CTS4386, CTS4406, CTS4489, CTS4535, CTS4687, CTS4912/Z1013, CTS5333/Z1014, CTS5343, CTS5444, CTS5451, CTS5477, CTS5920/Z1016, CTS5938, CTS6044, CTS6330, CTS6562, CTS6881, CTS6931, CTS6951, CTS7139, CTS7209, CTS7215, CTS7360, CTS7487, CTS7550, CTS7684, CTS7742, CTS7804, CTS7841, CTS7864, CTS8040, CTS8138, CTS8672, CTS8850, CTS8923/Z1019, CTS9016/Z1021, CTS9041, CTS9271, CTS9295, CTS9348, CTS9445, CTS9452, CTS9457, CTS9542, CTS9554, CTS9568, CTS9631/Z1022, CTS9814, CTS9847, CTS10142, CTS10204, CTS10246/Z1023, CTS10377, CTS10453, CTS10505, CTS10542, CTS10595, CTS10680, CTS10681, CTS10698/Z1029, CTS10968/Z1030, CTS11049/Z1031, CTS11086/Z1032, CTS11130, CTS11164/Z1033, CTS11207, CTS11226, CTS11254, CTS11318, CTS11472/Z1035, CTS11524/Z1036, CTS11817/Z1037, CTS11867, CTS11944, CTS12035, CTS12469, CTS12818, CTS12892, CTS13031, M234, SK973/Z15842, SK974/Z15797, Z994, Z998, Z1000, Z1002, Z1004, Z1006, Z1018, Z1025, Z1026, Z1038, Z1039, Z1040, Z15768, Z15769, Z15770, Z15771, Z15772, Z15773, Z15774, Z15775, Z15776, Z15777, Z15778, Z15779, Z15780, Z15781, Z15782, Z15783, Z15784, Z15785, Z15786, Z15787, Z15788, Z15789, Z15790, Z15791, Z15792, Z15793, Z15794, Z15795, Z15796.1/Z17076.1, Z15798, Z15799, Z15800, Z15801, Z15802, Z15803, Z15804, Z15805, Z15806, Z15807, Z15808, Z15809, Z15810, Z15811, Z15812, Z15813, Z15814, Z15815, Z15816, Z15817, Z15818, Z15819, Z15820, Z15821, Z15822, Z15823, Z15824, Z15825, Z15826, Z15827, Z15828, Z15829, Z15830, Z15831, Z15832, Z15833, Z15834, Z15835, Z15836, Z15837, Z15838, Z15839, Z15840, Z15841, Z15843, Z15844, Z15845, Z15846, Z15847, Z15848, Z15849, Z15850, Z15851, Z15852, Z15853, Z15854, Z15855, Z15856, Z15857, Z15858, Z15859, Z15860, Z15861, Z15862, Z15863, Z15864, Z15865, Z15867, Z15868, Z15869, Z15870, Z15871, Z15872, Z15873, Z15874, Z15875, Z15876, Z15877, Z15878, Z15880, Z15881, Z15882, Z15883, Z15884, Z15885, Z15887, Z15888, Z15889, Z15890, Z15891, Z15892, Z15893, Z15894, Z15895, Z15896, Z15897
• • • E2b1   M54, M90
• • • • E2b1a   M85
• • • •  E2b1a1   M200, CTS155/Z971, CTS462, CTS694, CTS775, CTS845, CTS893, CTS1480, CTS1611/Z980, CTS1618, CTS2768/Z981, CTS3109, CTS3136, CTS3470, CTS3583, CTS4171, CTS4257, CTS4992, CTS5832, CTS5991/Z984, CTS6510/Z1017, CTS7210, CTS8248, CTS8392, CTS8409, CTS8578, CTS8885
• • • •  E2b1a1a   P45
• • • •  E2b1a1b   P258
• • • E2b2   CTS1048, CTS1307, CTS1441, CTS1779, CTS3562, CTS4066, CTS5349, CTS5602, CTS5769, CTS6168, CTS6457, CTS6826, CTS9613, CTS10529, CTS11436, Z15898, Z15899, Z15900, Z15901, Z15902, Z15903, Z15904, Z15905, Z15906, Z15907, Z15908, Z15909, Z15910, Z15911, Z15912, Z15913, Z15914, Z15915, Z15916, Z15917, Z15918, Z15919, Z15920, Z15921, Z15922, Z15923, Z15924, Z15925, Z15926, Z15927

Private SNPs - After having been investigated, these SNPs have not met the population distribution criteria for placement on the tree. Either too few confirmed positive testers have been found OR multiple confirmed testers were confined to either a single surname or to a small group of related males.

SNPs under Investigation - Additional testing is needed to confirm adequate positive samples and/or correct placement on the tree.


Y-DNA haplogroup E would appear to have arisen in northern hemisphere of Africa based on the concentration and variety of E subclades in that area today. E1b1 is by far the lineage of greatest geographical distribution. It has two major sub-lineages, E1b1a and E1b1b. E1b1a is a lineage that originated and expanded from West or Central Africa to Eastern Africa and Southern Africa. E1b1b probably evolved either in Northeast Africa or the Near East and then expanded to the west--both north and south of the Mediterranean Sea. E1b1b1 clusters are seen today in Western Europe, Southeast Europe, the Near East, Northeast Africa and Northwest Africa. E2 is possibly of Central African origin and is spread in low frequencies across the continent. The Cruciani articles (references and links below) are indispensable resources for understanding the structure of this complicated haplogroup, but note that the Cruciani haplogroup labels are now superseded because of the recently discovered new SNPS that lie closer to the root of the E branch of the Y-haplogroup Tree.

A caution on clade labels: Because knowledge of this branch of the Y-chromosome tree has advanced so quickly in the last few years, different clade labels can be found in current use for the same SNP-determined branch of the tree. For example, it is still common to see E3b1 and E3b2 used to distinguish between the M78 and M81 branches of the tree though greater resolution is now possible.


Adams et al, The Genetic Legacy of Religious Diversity and Intolerance: Paternal Lineages of Christians, Jews, and Muslims in the Iberian Peninsula, American Journal of Human Genetics, 83(6): 725-36, 2008.
Alonso et al, The Place of the Basques in the European Y-chromosome Diversity Landscape. (available by subscription) European Journal of Human Genetics, 13:1293-1302, 2005.
Battaglia et al, Y-chromosomal Evidence of the Cultural Diffusion of Agriculture in Southeast Europe European Journal of Human Genetics, 249. 2008.
Behar et al, Contrasting Patterns of Y Chromosome Variation in Ashkenazi Jewish and Host Non-Jewish European Populations. (pdf) Hum Genet 114:354-365, 2004.
Bortolini et al, Y-Chromosome Evidence for Differing Ancient Demographic Histories in the Americas. American Journal of Human Genetics, 73:524-539, (2003).
Bosch et al, Paternal and Maternal Lineages in the Balkans Show a Homogeneous Landscape over Linguistis Barriers except for the Isolated Aromuns. Annals of Human Genetics, 70:459-87, (2006).
Capelli et al, Population Structure in the Mediterranean Basin: A Y Chromosome Perspective. (pdf) Annals of Human Genetics, 2005.
Cadenas et al, Y-chromosome Diversity Characterizes the Gulf of Oman. European Journal of Human Genetics, 16:374-386, 2008.
Cinnioglu et al, Excavating Y-chromosome Haplotype Strata in Anatolia. (pdf) Human Genetics. 114:127-148, 2004.
Cruciani et al, A Back Migration from Asia to Sub-Saharan Africa is Supported by High-Resolution Analysis of Human Y-Chromosome Haplotypes. American Journal of Human Genetics, 70:1197-1214, 2002.
Cruciani et al, Phylogeographic Analysis of Haplogroup E3b (E-M215) Y Chromosomes Reveals Multiple Migratory Events Within and Out of Africa. (pdf) American Journal of Human Genetics, 74:1014-1022, 2004.
Cruciani et al, Molecular Dissection of the Y Chromosome Haplogroup E-M78 (E3b1a): A Posteriori Evaluation of a Microsatellite-Networked-Based Approach Through Six New Biallelic Markers. (pdf) Human Mutation: Mutation in Brief #916, 2006.
Cruciani et al, Tracing Past Human Male Movements in Northern/Eastern Africa and Western Eurasia: New Clues from Y-Chromosomal Haplogroups E-M78 and J-M12. (pdf) Molecular Biology and Evolution 24(6):1300-1311, 2007.
Cruciani et al, Recurrent Mutation in SNPs within Y chromosome E3b (E-M215) Haplogroup: A Rebuttal. (abstract) American Journal of Human Biolology. Sep-Oct;20(5):614-6, 2008.
Deng et al, Evolution and Migration History of the Chinese Population Inferred from the Chinese Y-chromosome Evidence. (pdf) Journal of Human Genetics, 49:339-348, 2004.
El Sibai et al, Geographical Structure of the Y-Chromosomal Genetic Landscape of the Levant: A Coastal Inland Contrast, Annals of Human Genetics, 73:568-81, 2009. (abstract)
Firasat et al, Y-chromosomal Evidence for a Limited Greek Contribution to the Pathan Population of Pakistan, European Journal Human Genetics, 15:121+126, 2007.
Flores et al, Reduced Genetic Structure of the Iberian Peninsula Revealed by Y-chromosome Analysis: Implications for Population Demography. (pdf) European Journal of Human Genetics, 12:855-863, 2004.
Francalacci et al, Low-Pass DNA Sequencing of 1200 Sardinians Reconstructs European Y-Chromosome Phylogeny. Science: Vol. 341 no. 6145, pp. 565-569, DOI: 10.1126/science.1237947, 2 August 2013.
Gebremeskel et al, Y-chromosome E haplogroups: Their Distribution and Implication to the Origin of Afro-Asiatic Languages and Pastoralism. European Journal of Human Genetics; doi: 10.1038/ejhg.2014.41, 2014.
Gomes et al, Digging Deeper into East African Human Y Chromosone Lineages, Human Genetics, 127(5):603-13, 2010.
Henn et al, Y-chromosomal Evidence of a Pastoralist Migration through Tanzania to Southern Africa. PNAS, vol. 105 no. 31 10693-10698, 2008.
Karafet et al, New Binary Polymorphisms Reshape and Increase Resolution of the Human Y-Chromosomal Haplogroup Tree. Abstract. Genome Research, published online April 2, 2008. Supplementary Material.
King et al, The Coming of the Greeks to Provence and Corsica: Y-Chromosome Models of Archaic Greek Colonization of the Western Mediterranean, BMC Evolutionary Biology, 11: 69, 2011.
King et al, Differential Y-chromosome Anatolian Influences on the Greek and Cretan Neolithic. (abstract) Annals of Human Genetics. 72:205-214, 2008.
Magoon et al, Generation of High-resolution a priori Y-chromosome Phylogenies Using "Next-generation" Sequencing Data bioRxiv, doi: 10.1101/000802, December 13, 2013.
Nasidze et al, MtDNA and Y-chromosome Variation in Kurdish Groups. (abstract) Annals of Human Genetics, 69:401-412, 2005.
Regueiro et al, Iran: Tricontinental Nexus for Y-Chromosome Driven Migration. (abstract) Human Heredity, Vol. 61, No 3, 132-143, 2006.
Repping et al, High mutation rates have driven extensive structural polymorphism among human Y chromosomes. Nature Genetics 38, 463 - 467, 2006.
Rozen et al, Remarkably Little Variation in Proteins Encoded by the Y Chromosome's Single-Copy Genes, Implying Effective Purifying Selection. American Journal of Human Genetics. 2009 December 11; 85(6): 923-928.
Schuster et al, Complete Khoisan and Bantu Genomes from Southern Africa. (abstract) Nature 463, 943-947, 18 February 2010.
Semino et al, Ethiopians and Khoisan Share the Deepest Clades of the Human Y-Chromosome Phylogeny. (pdf) American Journal of Human Genetics, 70:265-268, 2002.
Semino et al, Origin, Diffusion, and Differentiation of Y-Chromosome Haplogroups E and J: Inferences on the Neolithization of Europe and Later Migratory Events in the Mediterranean Area. (pdf) American Journal of Human Genetics, 74:1023-1034, 2004.
Sengupta et al, Polarity and Temporality of High Resolution Y-chromosome Distributions in India Identify Both Indigenous and Exogenous Expansions and Reveal Minor Genetic Influence of Central Asian Pastoralists. (pdf) American Journal of Human Genetics, 78:202-221, 2006.
Shen et al, Reconstruction of Patrilineages and Matrilineages of Samaritans and other Israeli Populations from Y-Chromosome and Mitochondrial DNA Sequence Variation. (pdf) Human Mutation, 24:248-260, 2004.
Sims et al, Sub-Populations Within the Major European and African Derived Haplogroups R1b3 and E3a Are Differentiated by Previously Phylogenetically Undefined Y-SNPs. Human Mutation: Mutation in Brief #940, Online, 2007.
Trombetta et al, A New Topology of the Human Y Chromosome Haplogroup E1b1 (E-P2) Revealed through the Use of Newly Characterized Binary Polymorphisms. PLoS ONE 6(1), Online, 2011.
Valone et al, Y SNP Typing of African-American and Caucasian Samples Using Allele-Specific Hybridization and Primer Extension. (pdf) Journal of Forensic Science, 49:4, July 2004.
Zhao et al, Presence of Three Different Paternal Lineages among North Indians: A Study of 560 Y Chromosomes. (abstract) Annals of Human Biology, 36(1):46-59, 2009.

Additional Resources:

ISOGG Wiki - What you need to know about Genetic Genealogy.
E1a - (M33, M132) Project.
Haplogroup E1b1a (E1b1a1-M2), Roberta Estes.
The African DNA Project (E1b1a), Dr. Ana Oquendo Pabon.
Arabian E Y-DNA Project (E1b1).
E-M35 Project (formerly the E3b Project) (E1b1b1), Bill Harvey, Denis Savard, Victor Villarreal.
E-M35 Project - New SNP Tracker.
The E-M35 Phylogeny Project, Victor Villareal.
Double Helix Forums, E-M35 community.

Corrections/Additions made since 1 January 2015:

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