Y-DNA Haplogroup O and its
Subclades - 2017
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Because of
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Ray Banks
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^ Indicates
a next-generation sequencing
entry which does not yet meet quality guidelines for minimum number of
reads.
The
criteria for a representative SNP printed
in bold for a subclade is: traditional usage,
testing one
in multiple labs, and/or being found in the area of the chromosome used
in recent research studies.
SNPs
listed below in italics
(colored black or red)
are quality variants from next-generation sequencing reports
consistently showing as representing that subgroup.
Contact
Person for Haplogroup O: Owen Lu
O M175 ,
CTS647, CTS874/M1530, CTS1182/M1740, CTS1658/M1746,
CTS1853/M1747, CTS2340, CTS2419/M1748, CTS3283/M1749, CTS3771/M1750,
CTS5118/M1752, CTS5493/M1753, CTS7498/M1758, CTS7553/M1759,
CTS8384/M1762, CTS8746/M1763, CTS9039/M1766, CTS9544/M1767, CTS9577,
CTS9602/M1768, CTS10784/M1779, CTS10995/M1780, CTS11955/M1781,
F14/M1738, F45/M1739, F84/M1741, F137/M1743, F171/M1573, F175,
F182/M1744, F282/M1359, F315/M1751, F355/M1754, F371/M1756,
F380/M1757/S27659, F452/M1761, F478/M1764, F494/M1765, F537/M1769,
F540, F546/M1770, F600/M1777, F614/M1778, F668/M1782, FGC12433,
FGC12434, FCG12436, L1361/M1771, M1742, M1745,
M1772, M1773, M1774, M1775, M1776 , P186, P188, P191, P196, Z16962,
Z16966 O1 F265/M1354 ,
CTS2866, F75/M1297, F429/M1415, F465/M1422 O1a M119 ,
CTS31, F589/Page20, L246,
L466 O1a1 B384/Z23193 O1a1a M307.1/P203.1 ,
CTS3422, CTS4351, CTS5059,CTS6864, CTS7015, CTS8229, CTS8875, CTS8934,
CTS9321, CTS11688, F31, F54, F89, F303, F333, L83 O1a1a1 F446 ,
CTS4588, F560, FGC15381/K620/Z23389, K613/Z23387, K619/Z23388
O1a1a1a F140 ,
CTS611, CTS3089, CTS3265, CTS3269,CTS11270, F157,
F343, F424, F518, F571, FGC15382/Z23466,
FGC15391/Y14275/Z23470, FGC15383/K625/Z23474, V68.2, Z23457 O1a1a1a1 F78 ,
CTS3758, CTS10963, F168, F206
O1a1a1a1a F81
O1a1a1a1a1 F533
O1a1a1a1a1~ CTS2458 O1a1a1a1a1a F492,
CTS2498, CTS2594, CTS4206,CTS5075, CTS11784^, F619,
K629/Z23478 O1a1a1a1a1a1 F656 ,
Z23481
O1a1a1a1a1a2~ CTS409, CTS2245, CTS4924, CTS5922,
CTS11553, K630/Z23504, Z23494, Z23495, Z23496, Z23499, Z23505
O1a1a1a1a1b CTS4585,
FGC15395/Y14269, FGC15397/Y14272, FGC15398/Y13987,
FGC15399/Y13986, FGC15400/Y13989, FGC15401/Y13988, FGC15402/Y13984,
FGC15409/Y13985, Y14273 O1a1a1a2 YP4610/Z39229 O1a1a1a2a AM00330/AMM480/B386 ,
SK1533, Z39230/YP4600, Z39231/YP4601, Z39232/YP4603, Z39233/YP4605,
Z39234/YP4607, Z39235/YP4609, Z39236/YP4611, Z39237/YP4612,
Z39238/YP4613, Z39239/YP4614, Z39240/YP4615, Z39241/YP4616,
Z39242/YP4617, Z39243/YP4618, Z39244/YP4619, Z39245/YP4620,
Z39246/YP4621
O1a1a1a2a1 AM00333/AMM483/B387,
Z39247/YP4602
O1a1a1a2a1a B388,
Z39248/YP4604, Z39249/YP4608
O1a1a1a2b SK1555 O1a1a1b SK1568/Z23420 O1a1a1b1 M101 O1a1a1b2 Z23392 ,
Z23404, Z23440 O1a1a1b2a Z23442
O1a1a1b2a1 SK1571 ,
Z39268
O1a1a2 CTS52 ,
CTS5880, CTS8423 , CTS11785, F4084
O1a1a2a CTS701 ,
CTS4115, CTS6217 O1a1a2a1 K644/Z23266 ,
Z23269 O1a1b CTS5726 ,
CTS3085, CTS3400, CTS7370^,CTS8228^, CTS9138^, CTS9243^, CTS9540^,
CTS11079, F819^, F1452, F1620.2, F2104, F2644^, F2852^, F3063,
F3495,F3640, SK1589, Z39164, Z39165, Z39166^, Z39167, Z39168, Z39169,
Z39170, Z39171^, Z39172^, Z39173, Z39174,Z39175^, Z39176, Z39177,
Z39178, Z39179^, Z39180^, Z39181, Z39182, Z39183^, Z39184, Z39185,
Z39186, Z39187,Z39188, Z39189, Z39190, Z39191, Z39192, Z39193, Z39194,
Z39195, Z39196^, Z39197, Z39198^, Z39199^ O1a1b~ F2911 O1a2 M110 ,
F881, F908, F1081,F1608, F1880, F2340, F2380, F2635, F2671, F3134,
F3152, F3157, F3214, M50, M103 O1a2a F3288 O1a2a1 B392 ,
Z38606,Z38607, Z38608, Z38609, Z38610, Z38611 O1a2a1a B393 ,
Z38612 O1a3~ Page109 O1b M268 ,
F138, F167/L690, F256/M1341, F292/M1363, F330/L463, F435/M1417, F516, P31 O1b1 F2320 ,
F3008/M1441, F3286/M1465 O1b1a M1470 O1b1a1 PK4 ,
CTS6616/M1392, CTS7320/M1405, CTS7541/M1407, CTS7548/M1408,
CTS8589/M1425, CTS9150, CTS11303, F722/M1279, F932/M1290, F1058/M1302,
F1085/M1305, F1130/M1306, F1265.1/M1313.1, F1346/M1316, F1353/M1317,
F1809/M1352, F2317, F2338.2/M1391.2, F3033.1/M1443.1/V4056.1/Z2838.1,
F3424/M1489, F3439/M1494 O1b1a1a M95 ,
F709, F987/M1300, F1931/M1362, F2176/M1376, F2254/M1383, F2917/M1436,
F4233/M1404
O1b1a1a1 F1803/M1348 ,
CTS6251/M1384, M1310
O1b1a1a1a F1252
O1b1a1a1a1 F2924 ,
CTS10077 , CTS10240
O1b1a1a1a1a M111 ,
M88, CTS1163, F1120, F1643, F2346, F2454, F2524, F2554, F2861, F3055
O1b1a1a1a1a1 F2758 ,
CTS10965, Z24081,
Z24082, Z24084, Z24085, Z24087
O1b1a1a1a1a1a Z24083 ,
Z24086
O1b1a1a1a1a1a1 Z24089
O1b1a1a1a1a1a1a F923 ,
F1108, F2411, F3671
O1b1a1a1a1a1a1a1 CTS2022
O1b1a1a1a1a1a1a1a F1399 ,
F3053
O1b1a1a1a1a1a1a1a1 F2415 ,
CTS3745, Z3955.1
O1b1a1a1a1a1a1a2 Z24131
O1b1a1a1a1a1a1a3 Z24100 O1b1a1a1a1a1a1b SK1627/Z24091 ,
Z24093
O1b1a1a1a1a1a1b1 Z39410
O1b1a1a1a1a1a2 Z24088
O1b1a1a1a1a2 F2890
O1b1a1a1a1a2a Z24048
O1b1a1a1a1a2a1 Z24050
O1b1a1a1a1a2b Z24014 O1b1a1a1a1b CTS5854 ,
Z23743, Z23744 O1b1a1a1a1b1 Z23810
O1b1a1a1a1b1a CTS7399 ,
FGC19685, FGC19699
FGC19670/Y14253/Z23825, FGC19672/Y14260/Z23840,
FGC19673/SK1620/Y14000/Z23821, FGC19674/Y14018/Z23835,
FGC19675/Y13998/Z23819, FGC19676/Y14255/Z23831, FGC19677/Y14259/Z23834,
FGC19678/Y14257, FGC19679/Y14008/Z23828, FGC19681/Y14020/Z23836,
FGC19682/Y14261/Z23841, FGC19683/Y14011/Z23829, FGC19684/Y14014/Z23832,
FGC19686/Y14022/Z23839, FGC19687/Y13999/Z23820, FGC19688/Y14007/Z23827,
FGC19689/SK1619/Y14001/Z23822, FGC19694/SK1618/Y14256, FGC19705/Y14010,
FGC19710/Y14021/Z23837, Z23824, Z23833, Z23838, Z23843
O1b1a1a1a1b1a1 FGC19713/Y14026 ,
FGC19671/Z23845,
FGC19690/Y14251/Z23851, FGC19693/Y14013/Z23857, FGC19698/Y13996/Z23852,
FGC19700/Y14017/Z23861, FGC19701/Y14002/Z23853, FGC19703/Y14262/Z23866,
FGC19711/Y14025/Z23865, FGC19712/Y14016/Z23860, FGC19715/Y13997, FGC19724 , Z23858
O1b1a1a1a1b1a1a Z23849
O1b1a1a1a1b1a1a1 FGC61038 , FGC61039, FGC61054, Y31276, Y31927, Y31928
O1b1a1a1a1b1b CTS651 ,
CTS5267, CTS7642, CTS8722, CTS9915 O1b1a1a1a1b1b1 CTS9884 O1b1a1a1a1b2 F4229 ,
F4212, F4251, FGC61079, FGC61080, PF3452, Z23746, Z23769, Z23781, Z23787, Z23799 O1b1a1a1a1b2a F809 , Y31259, Y32109, Z23745, Z23766, Z23777, Z23784, Z23786, Z23791, Z23797, Z23801, Z23803, Z23804, Z23807 O1b1a1a1a1b2a1 F2517, Z23795
O1b1a1a1a2 SK1630 ,
SK1631, SK1632, SK1633 O1b1a1a1a2a SK1636
O1b1a1a1b F789/M1283 ,
F2028/M1367,F2522/M1410, F3092, F3318/M1469, F4181/M1357
O1b1a1a1b1 FGC29900/Y9322/Z23667 O1b1a1a1b1a B426/FGC29896/Y9033/Z23671 ,
FGC29897/Y9320/Z23673, FGC29898/Y9036/Z23674, FGC29899/Y9034/Z23672
O1b1a1a1b1a1 FGC29907/YP3930 ,
SK1640/YP3931 O1b1a1a1b1a2 B427/Z23680 ,
M6661 O1b1a1a1b1b Z39485
, Z39486^, Z39487, Z39488, Z39489^ O1b1a1a1b1c B418 ,
Z39490
O1b1a1a1b2 SK1646
O1b1a1a2 CTS350 ,
CTS2549, CTS3471,CTS6531, CTS7187, CTS7270, CTS7809, CTS11962.2/Z2953.2
O1b1a1a3~ Page103 O1b1a1b F838 ,
CTS892 O1b1a1b1 F1199 ,
F1309, F1685,F1742, F2189, F2445, F2995, F3353 O1b1a2 Page59 ,
CTS1456, CTS1719, CTS4040, CTS5528, CTS5745, CTS7643, F883, F888,
F1098, F1485, F2117, F2290, F2601, F3016, F3079, F3445 O1b1a2a F993 ,
CTS2335, CTS5445,CTS7866, CTS8849, CTS10154, CTS10437, CTS11033, F1484,
F1669, F2120, F2124, F2131, F2383, F2414, F2590, F2771,F2862, F3036 ,
F3135, F3217, F3262, SK1659, Z24391 O1b1a2a1 F1759 ,
CTS5847,CTS8414, CTS11890, F2064, F3314, F3323, F3346, F3478, Z24393,
Z24407
O1b1a2a1a CTS1127 A16611, A16612, A16613, CTS6105, PH2506, PH2797, PH5023 O1b1a2b F417/M1654 ,
CTS7268, CTS10113,F1236, F1860, F1956, F2061, F2504, F2775, F3252,
F3291, F3632 O1b1a2b1 F840 ,
CTS250, CTS949, F984, FGC30073, Y10774
O1b1a2b1a F1127 ,
CTS10685, F1628, F1776, F2183
O1b1a2b2 CTS1451 ,
CTS4475, CTS5284, CTS6503, CTS9341 O1b1a2c CTS9996 O1b2 P49 ,
M176/Page63/SRY465 O1b2a F1942/Page92 ,
IMS-JST022454, M302 O1b2a1 CTS9259 ,
CTS348, CTS9752, F2315, PF4474, Z24512, Z24520, Z24522, Z24553
O1b2a1a F1204 ,
F1290, F1530, F1805,F2634, F2960, F3147, F3327, F3356, K10 O1b2a1a1 CTS713 ,
CTS7776, CTS8032, CTS10145, CTS10341, CTS10674, CTS11723, CTS11986, K7
O1b2a1a1a CTS1875 ,
CTS7465, CTS8777
O1b2a1a1a1 CTS10682 ,
CTS1966,CTS3764.2, CTS3926, CTS6725, CTS7366, CTS8034, Z24596
O1b2a1a1b Z24598 ,
Z24599
O1b2a1a1c CTS203 ,
CTS6622, CTS9892
O1b2a1a2 F2868 ,
F3110, K4
O1b2a1a2a L682 ,
CTS2734, CTS6803,CTS9322, K28
O1b2a1a2a1 CTS723 ,
Z24624
O1b2a1a2a1a CTS7620 ,
Z39506
O1b2a1a2a1b A12446 , A12447,
A12448
O1b2a1a2a1b1 PH40 , PH437
O1b2a1a2b F940 ,
F1912, F3390
O1b2a1a3 CTS10687 ,
F1800, F1813, F1823, F1877, F3821
O1b2a1a3a CTS1215 ,
CTS3729, F787, F867, F1553, F2026, F2604, F2767, F3065
O1b2a1b CTS562 ,
CTS3505, CTS10354 O1b2a2~ Page90 O2 M122 , CTS10736, CTS10753, F36, F400, F433, F471,
F633,P198 O2a M324 ,
CTS8153, CTS8399, F27, F71, F113, F129, F166, F341, F406, F572,
P93/Page79, P197, P199, P200 O2a1 L127.1 ,
CTS5687, CTS7638,CTS11981, F51, F58, F95, F145, F159, F164, F170, F243,
F272, F300, F304, F305, F412, F498, F573, F601, F642, F676, KL1/L465,
KL2/L467 O2a1a F1876/Page127 ,
F964, F3143
O2a1a1 F2159, F2726, F2789, F3169, F4102 O2a1a1a F1867/Page124 O2a1a1a1 F852, CTS3709 , CTS8168, CTS8587, CTS10730, F1054, F4348/Z24834, SK1669/Z24838, Z24836, Z24840, Z24844
O2a1a1a1a F2266
O2a1a1a1a1 L599
O2a1a1a1a1a Z43961 , Z43962
O2a1a1a1a1a1 Z43963 , Z43964, Z43965
O2a1a1a1b F854
O2a1a1a1b1 Z43966, Z43967, Z43968, Z43969, Z43970, Z43971, Z43972, Z43973, Z43974
O2a1a1a1c~ Page130
O2a1a1b F915 ,
CTS7667, F1478, F1675, F1920, F2528, F2633, F2975, F3046, F3182, F3284,
F3287, F3417, F3656, F3669, F4195, F4312, PF5390 O2a1a1b1 CTS1936 ,
Z24782, Z24797, Z24818
O2a1a1b1a Z43975 , Z43976, Z43977, Z43978, Z43979, Z43980 O2a1a1b2 FGC33994 ,
FGC34004, FGC34008, FGC34009, FGC34019, FGC34022, FGC34023, FGC34029 O2a1b~ M164 O2a1c IMS-JST002611 ,
CTS1872, CTS2483, CTS5879, F61, F99, F240, F247, F302, F309, F460, F562 O2a1c1 F18 ,
CTS11342
O2a1c1a F117 , CTS5820, F514, FGC12471/Y3268
O2a1c1a1 F13 , F30, F62 , F68, F251 , F253, F379 , F530
O2a1c1a1a F11 ,
F60, F325, F362 ,
F526, F561 ,
F425/Page69, FGC12472/Y3271/Z25059
O2a1c1a1a1 F632 ,
CTS4580, Z25061
O2a1c1a1a1a F110/M11115 ,
CTS10716,
F133,
F434, F646, Y21437, Y21448 , Z25064, Z25065,
Z25067, Z25068
O2a1c1a1a1a1 F17 ,
CTS362, CTS2801 , CTS3663, CTS4414, CTS5822, CTS6777, CTS9127 , CTS11839, F64, F92 , F357, F529, F620, F672, F691, Z25075, Z25076, Z25077, Z25079, Z25081, Z43887
O2a1c1a1a1a1a F377
O2a1c1a1a1a1a1 F1095 ,
CTS7789, F1249, F1250,
O2a1c1a1a1a1a1a F856
O2a1c1a1a1a1a1a1 F1418 , F1495
O2a1c1a1a1a1a1a2 Z25097 , Z25098
O2a1c1a1a1a1a2 CTS7501
O2a1c1a1a1a1b F793 ,
F1316C , F2035, F2108, F3433, Z43869, Z43870, Z43871, Z43872, Z43873, Z43874, Z43875, Z43876, Z43877, Z43878
O2a1c1a1a1a2 Y20951 , Z43879
O2a1c1a1a1a2a Y20932 ,
Y20933, Y20934, Y20935, Y20936, Y20937, Y20938, Y20939, Y20940, Y20941,
Y20942,Y20943, Y20944^, Y20945, Y20946, Y20947, Y20948^, Y20949,
Y20950^, Y21435, Y21438, Y21439, Y21442,
Y21443, Y21444, Y21445, Y21446, Y21447 , Z34692, Z43880. Z43881, Z43882, Z43883, Z43884, Z43885, Z43886
O2a1c1a1a2 F38 ,
F106, F136, F142, F178, F270, F286, F358, F381,
F386, F475, F479, F485, F553, F592, F625, F1978,
F3131, F3453
O2a1c1a1a3 F12 ,
F196, F480, F659, F660
O2a1c1a1a4 F930 ,
F1232, F2356, F2589
O2a1c1a1a4a F2685
O2a1c1a1a5 F1365/M5420/PF1558 ,
F921 , F1676, F2109, F2180, F2213, F3232, F3452,
F4062, FGC54466,
FGC54513, FGC54523, FGC54527 , Y15975, Y15977 , Y15978,
Y15979, Y16155
O2a1c1a1a5a Y15976 ,
Y15980,
Y26385
O2a1c1a1a5a1 Y16154
O2a1c1a1a5a1a Y26383 , Y26387,
Y27899
O2a1c1a1a5a1a1 SK1686 FGC49730,
Y26384, Y26386, Y27894, Y27895, Y27896, Y27897, Y27898, Y27900
O2a1c1a1a5b FGC54486 , FGC54495,
FGC54520, FGC54522, FGC54529, FGC54538, FGC54540, FGC54545, FGC54488
O2a1c1a1a5b1 FGC54507
O2a1c1a1a6 CTS12877 ,
Z25165
O2a1c1a1a6a F2527
O2a1c1a1a6a1 CTS5409 ,
CTS6236
O2a1c1a1a6a2 F2941 ,
F4073, F4119
O2a1c1a1a7 F723 ,
F971, F972, F1210, F1351, F1422, F1638, F2357, F2560, F2719, F3042,
F3103, F3270, F4171
O2a1c1a1a8 CTS2107 ,
CTS2128, CTS2916, CTS3518, CTS4150, CTS8491, CTS10670
O2a1c1a1a9 SK1691 , SK1692,
SK1695, SK1696
O2a1c1a1b PH203 ,
FGC1803.2/Y2044.2, PH278, PH975, PH1787, PH2385, PH3699, PH3838,
PH3849, PH4404, PH4535, PH4997, PH5031, PH5323 O2a1c1b F449
O2a1c1b1 F238
O2a1c1b1a F134 ,
F271, F322
O2a1c1b1a1 F1273 , F1894, F2468, F3256, F3309, F3434
O2a1c1b1a2 F724 ,
F1156, F1336, F1910 O2a1c1b2 F1266 ,
F1497, F2016, F4267 O2a1c1c CTS498 , CTS957, CTS1028, CTS2398, CTS4941, CTS5926, CTS7341, CTS8263, CTS9516, CTS10139, CTS10185, CTS11076, CTS11949 O2a1c2 FGC3750/SK1673 ,
L133.2/Page74.2 O2a2 IMS-JST021354/P201 ,
CTS8236, F525 O2a2a M188
O2a2a1 F2588 ,
CTS3106, F2638
O2a2a1a CTS445
, CTS11727, F2055, F2221, F2329, F2592, F3559
O2a2a1a1 CTS201
O2a2a1a1a M159/Page96
O2a2a1a2 M7 ,
CTS10944, CTS12052 , F2591, F2594, F2680, F3177, F3342, F3371, F4066, M209, Z25245
O2a2a1a2a F1276 ,
F1898, F2179, F2428, F2962, F3406, F4064, F4173
O2a2a1a2a1 CTS6489
O2a2a1a2a1a F1275 ,
F2309, F3085
O2a2a1a2a1a1 M113
O2a2a1a2a1a2 N5 ,
F1100, F1234, F1411, F3229, F3516
O2a2a1a2a1a3 Z25400
O2a2a1a2a2 F1863 ,
F2079
O2a2a1a2a2a F1134 ,
F1992, F2332, F2480, F2612
O2a2a1a2a2a1 F1262 ,
F2208, F2515, F3469
O2a2a1a2b Y26403 ,
Y26409, Y26412, Y26413, Y26415, Y26416, Y26421, Y26422, Y26424, Y26428,
Y26430, Y26433, Y26434, Y26437, Y26440, Y26446, Y26448, Y26457, Y26460,
Y27924, Y27927, Y27932, Y27933, Y27934, Y27939, Y27942, Y27943, Y27951, ZS2079
O2a2a1b F1837
, F2576, F2928, F4088, F4163 O2a2a2 F879
O2a2a2a F1226
O2a2a2a1 F2859 ,
F2937 O2a2b P164 ,
CTS4723, CTS11109, CTS12099, F130, F131, F299, F422, F427 O2a2b1 M134 ,
F100/M1546
O2a2b1a F450/M1667
O2a2b1a1 M117,
Page23 , CTS899/M1531, CTS1275/M1536, CTS3251, CTS5128/M1619,
CTS6623/M1638, CTS11742/M1720, F141/M1564, F144, F235/M1587,
F342/M1627, F373/M1636, F476/M1671, F579/M1692, F581, F584, F613/M1702,
F649
O2a2b1a1a M133 ,
F8, F42
O2a2b1a1a1 F438
O2a2b1a1a1a Y17728
O2a2b1a1a1a1 F155 ,
CTS630, CTS1304, CTS2760,
CTS11316 ,
F201, F254, F276, F316, F414, F440, F541,
F703, F6227, F6517, F6530, F6872, F6909^, F7446, F7447, F7556, F7973,
F7994, F14281, F14428, F14445, F14663^, Y21365, Y21366, Y21367
O2a2b1a1a1a1a F813/M6539 ,
Y21364
O2a2b1a1a1a1a1 Y20928
O2a2b1a1a1a2 F1754
O2a2b1a1a1a2a F2137 , A16631, A16632 ,
F2503, F2643, S24099.2
O2a2b1a1a1a2a1 F1442 , F2779
O2a2b1a1a1a2a1a F1123 , F2265, F2425, F2453, F3048, F3115 O2a2b1a1a1a2a1a1~ F1369
O2a2b1a1a1a2a2 A16636 , A16637, A16638, A16639, A16640, A16641, A16642, A16643, A16644, A16645
O2a2b1a1a1a3 Z25907 ,
Z25908, Z25909
O2a2b1a1a2 FGC23469/Z25852
O2a2b1a1a2a F310 ,
F666, F692
O2a2b1a1a2a1 F402 ,
F624
O2a2b1a1a2a1a F1531 ,
F2781
O2a2b1a1a3 CTS7634 O2a2b1a1a3a F317 ,
F375, F474, F574, F869, F1401, F1707, F1870, F2000, F2188, F2494,
F2631, F2845, F3212, F3431, F3455, Y16475, Y16476, Y16478, Y16479,
Y16696
O2a2b1a1a3a1 F3039
O2a2b1a1a3a2 Y29861 , Y30150
O2a2b1a1a3b CTS5488 ,
PH3164, PH3555, Z39594
O2a2b1a1a4 Z25853 O2a2b1a1a4a CTS5492 ,
CTS4789,
CTS5063, CTS7418, CTS8299, CTS1265 5
O2a2b1a1a4a1 CTS6987 ,
Z25902, Z25903, Z25905
O2a2b1a1a4a1a Z42620
O2a2b1a1a4a2 F20963 , Z44926, Z44927
O2a2b1a1a5 CTS10738/M1707 ,
CTS1017/M1532, CTS7316, M1543, M1694, M1726 O2a2b1a1a5a CTS9678 ,
CTS8848/M1672, CTS10804/M1708, CTS11844, M1540,
M1541, M1575, M1586, M1624, M1650, M1698, M1701, M1722, M1732, Z25850
O2a2b1a1a5a1 Z39663 ,
Z39664
O2a2b1a1a5a2 M1513 , M1557, M1593, M1602, M1604, M1608, M1617, M1629, M1637, M1683, M1688, M1719, Z44923, Z44924, Z44925
O2a2b1a1a5b A9457 ,
A9458 F16367, F24592
O2a2b1a1a5b1 F17158 , Z44918, Z44919, Z44921, Z44922
O2a2b1a1a6 CTS4658 ,
CTS1642, CTS9332
O2a2b1a1a6a CTS5308 O2a2b1a1a6b Z25928,
Z25929, Z25930, Z25931
O2a2b1a1a6b1 SK1730/Z25982 ,
Z25975, Z25981, Z25983, Z25984, Z25985, Z26000
O2a2b1a1a6b1a Z26030 ,
Z26031
O2a2b1a1a6b1b Z26010 ,
Z26012, Z26013, Z26014
O2a2b1a1a6b2 A9462 ,
A9461, A9463, A9464, A9465, A9466, A9467, A9468
O2a2b1a1a6b3 B456 ,
Z39703, Z39704, Z39705, Z39706, Z39707, Z39708
O2a2b1a1a7 YP4864 ,
YP4865, Z44052, Z44053, Z44054, Z44055, Z44056, Z44057, Z44058, Z44059,
Z44060, Z44061, Z44062, Z44063, Z44064, Z44065, Z44066, Z44067
O2a2b1a1a7a Z44068 , Z44069
O2a2b1a1a7a~ F5524/SK1746, Z44070
O2a2b1a1a7a1~ F5525/SK1748
O2a2b1a1a7b Z44071 , Z44072,
Z44073, Z44074, Z44075, Z44076, Z44077, Z44078, Z44079, Z44080, Z44081,
Z44082, Z44083, Z44084, Z44085, Z44086, Z44087, Z44088, Z44089, Z44090
O2a2b1a1a8 Z44091
O2a2b1a1a8a Z44092 , Z44093, Z44094
O2a2b1a1b CTS4960 ,
SK1717, SK1720, SK1721 O2a2b1a2 F114 ,
CTS7655, F122, F274/Page125, F403, F430, F444, F499/Y19, F593, F701
O2a2b1a2a F79 ,
F314, F372, F629
O2a2b1a2a1 F46/Y15 ,
CTS2643, CTS11192, CTS11459, F174, F323/Y17, F353, F388, F442, F502,
F507/Y18, F532, F2039 O2a2b1a2a1a FGC16847/Z26091
O2a2b1a2a1a1 F48 ,
CTS326 , CTS816, CTS1011, CTS3871, CTS11177 ,
F163, F275, F523, F575 O2a2b1a2a1a1a F152 ,
CTS1098, CTS8380, CTS9172, CTS9217, F279, F595,
F700 O2a2b1a2a1a1a1 F2505 O2a2b1a2a1a1b CTS3149 ,
CTS5323, CTS5443, CTS1255 1
O2a2b1a2a1a2 F242 ,
CTS6493, CTS10286, CTS10401, CTS10888,
CTS11492, F273, F563, F634, F1116 O2a2b1a2a1a2a CTS4266 O2a2b1a2a1a2a1 Z26108 ,
Z26109, Z26110, Z26111, Z26112, Z26113 O2a2b1a2a1a2a1a F2173 ,
F1458, F3387, F3580 , F3614
O2a2b1a2a1a3 F2887 ,
CTS3776, CTS9862
O2a2b1a2a1a3a F3607 O2a2b1a2a1a3a1 F3525 O2a2b1a2a1a3b CTS3763 ,
FGC16857
O2a2b1a2a1a3b1 A9472
O2a2b1a2a1a3b2 FGC16863/Y7110
O2a2b1a2a1a3b2a L1360 ,
FGC16864, FGC16866, FGC16867/Y13527, FGC16869/Y13528, FGC16948
O2a2b1a2a1a3b2a1 FGC16889 ,
FGC16891, FGC16913, FGC16946
O2a2b1a2a1a3b2b SK1768/Y7112/Z26257 ,
CTS2083, CTS2119, CTS7323, FGC23839/Y7114/Z26259,
FGC23841/Y7116/Z26268, Y7111/Z26256, Z26266 O2a2b1a2a1a3b2b1 F4249 ,
FGC1255, FGC23844, FGC23850, FGC23852, FGC23854, FGC23855,
FGC23856, FGC23857, FGC23858, FGC23860, FGC23862, FGC23863, FGC23864,
FGC23865, FGC23870, FGC23871, FGC23873, SK1769, SK1770, Y16776, Y16777,
Y16778, Y16779
O2a2b1a2a1a3b2b1a FGC23868
O2a2b1a2a1a3b2b2 CTS335 ,
CTS558, CTS2080.2, CTS2882, CTS3249, CTS5450, CTS7951, CTS8942,
CTS10883, CTS11454 O2a2b1a2a1b CTS53 ,
CTS448, CTS610, CTS4696, CTS10819, CTS11910, SK1756/Z26318
O2a2b1a2a1b1 CTS6373
O2a2b1a2a1b1a A9473 O2a2b1a2a1c F3386
O2a2b1a2a1d Y29828 , Y30477, Y30485, Y30487, Y30489, Y30493, Y30498, ZS11791
O2a2b1a2a1d1 F735 , F2903 , L932.2, Y29836, Y30471, Y30484, Y30499
O2a2b1a2a1d1a FGC34973 ,
Y29829, Y29830, Y29831, Y29832, Y29833, Y29834, Y29835, Y30465, Y30466,
Y30467, Y30468, Y30469, Y30470, Y30472, Y30473, Y30474, Y30475, Y30476,
Y30478, Y30479, Y30480, Y30481, Y30482, Y30483, Y30486, Y30488, Y30490, Y30491, Y30492, Y30494, Y30495, Y30496, Y30497, Y30500
O2a2b1a2a1d1b~ F1739 O2a2b1a2b F743 ,
CTS1933, CTS2260, CTS8043, CTS10219,
F1725, F14400/Z26072,
F14403/Z26073, F14487/Z26074, F14497/Z26075, F14503/Z26076,
F14559/Z26077, F14690/Z26080, F14695/Z26081, F14714/Z26083, FGC61160,
FGC61167, Y31256, Z26078
O2a2b1a2b1 CTS8481 , CTS2205, FGC61159, FGC61164, FGC61165, FGC61215, Y31257
O2a2b1a2b1a CTS4325 , A16615,
A16616, A16617, A16618, A16621, A16622, A16623, A16624, A16625, A16626,
A16628, CTS496, CTS724, CTS1707, CTS2147, CTS2390, CTS4336, CTS5529, CTS7277 , CTS8451, CTS9362, CTS11009
O2a2b1a2b1a1 A16629 , A16630
O2a2b1a2b1a2 CTS682 ,
CTS2272, CTS7184, CTS8951
O2a2b1a2b2 F748 ,
F823, F1448, F2093, F2095, F2212, F2236, F2530, F2795, F2974, F3279,
F3383, F3440, F3505, F3579 O2a2b1a2b2a F728 ,
F1060, F2018, F2326 O2a2b1a2c~ Page101 O2a2b2 AM01822 /F3223 , AM01747/F1672, AM01749/F1693,
AM01770/F2029, AM01776/F2083, AM01777/F2139, AM01788/F2469,
AM01800/F2683 , AM01823 /CTS1366/F3237,
AM01828/N6 , F996, F1481, F1598/ S3543
O2a2b2a AM01856 /F871
O2a2b2a1 N7 ,
F2472, F4079, F4286, F4328, F4329, F4333 Z25701, Z25708, Z25713, Z25716, Z25720, Z25721, Z25728, Z25730, Z25737, Z44816
O2a2b2a1a F4110 ,
F4144 CTS12480, F4215, F20220, F20394, F24749, Z44734, Z44735, Z44736, Z44737, Z44738, Z44739
O2a2b2a1a1 F4068 , F4139, F4159, F4190, F4202, F4241, F4276, F4293, F4332, F4335
O2a2b2a1a2 SK1780 , SK1781
O2a2b2a1b F4124 SK1778/Z25746,
Z25747, Z25748, Z25749, Z25750, Z25752, Z25754, Z25755, Z25756, Z25757,
Z25758, Z25759, Z25760, Z25761, Z25762, Z25763, Z25764, Z25765, Z25767
O2a2b2a1b1 IMS-JST008425p6 A16609, A16610, F24792, F26088, FGC14331.2/ZS4034.2, FGC28699, SK1777, Z8263, Z40733,
Z44740, Z44741, Z44742, Z44743, Z44744, Z44745, Z44746, Z44747, Z44748,
Z44749, Z44750, Z44751, Z44752, Z44753, Z44754, Z44755, Z44756, Z44757,
Z44758, Z44759, Z44760, Z44761, Z44762, Z44763, Z44764
O2a2b2a1b2 BY15188 ,
F14875, F15739, F15692, F15973, F16170, F17339, F18455, F18521, F18757,
F19055, F20227, F20235, F20251, F20767, F21091, F21275, F22909, F23173,
F23473, F23524, F23679, F24517, F24605, F25036, F25041, F25079, F25381,
F25411, F25440, F25789, F25924, F26193, F26484, F26752, Z44765, Z44766,
Z44767, Z44768, Z44769, Z44770, Z44771, Z44772, Z44773, Z44774, Z44775,
Z44776, Z44777, Z44778, Z44779, Z44780, Z44781, Z44782, Z44783, Z44784,
Z44785, Z44786, Z44787
O2a2b2a1b2a F16411 , F17835, F19270, F19578, F22244, F22838, F23276, F23329, F24971, Z44792
O2a2b2a2 AM01845 /F706 ,
AM01755, AM01758/F1903,
AM01768, AM01780, AM01790/F2486, AM01791, AM01792, AM01799/F2654, AM01801, AM01804, AM01814/F3069 AM01842, AM01857, F1645, F16175, Z44817 O2a2b2a2a F717 ,
F919, F976, F999F1010, F1375, F1412, F1884, F1889, F2132, F2511, F2558. F2724, F2929, F3066, F3095, F3230, F3473, F3820, SK1782, SK1784 O2a2b2a2a1 F3612 , F3819 O2a2b2a2a2 SK1783 O2a2b2a2b AM01847/B451 , AM01750, AM01759, AM01760, AM01761/SK1787, AM01762, AM01763, AM01771, AM01775, AM01778, AM01781/S18967/SK1786, AM01785, AM01786, AM01787, AM01794, AM01798/SK1788 AM01802,
AM01805, AM01806, AM01812, AM01813, AM01815, AM01818, AM01821, AM01824,
AM01827, AM01835, AM01838, AM01839, AM01844, AM01855, AM01859, AM01860,
F15980, F16102, F16868, F17803 O2a2b2a2b1 A 17418 ,
A17419, A17420, A17422, A17424, A17425, A17426, A17427, A17428, A17429,
A17430, A17431, A17432, A17433, A17434, A17436, A17437, A17438, A17439,
A17441, A17442, A17443, A17444, A17445, A17446, A17447, A17448, A17449,
A17450, A17451, YP6194 O2a2b2a2b2 AM01756 , AM01757, AM01779, AM01803, AM01810, AM01826, AM01834, AM01861, F16663, F16771, F16783 O2a2b2a2b2a B450 , AM00474/SK1791, AM00475/SK1792, AM00476, F16769 O2a2b2a2b2b AM00472/B452 , AM00463, AM00464, AM00465, AM00466, AM00467, AM00468, AM00469, AM00470, AM00471, F16088, F16616, F16997, F17764, F26682 O2a2b2a2b2b1 F18942 , F21615, F25531 O2a2b2a2b2c A16427 , A16428, A16429, A16430, A16431, A16432
O2a2b2b A16433 , A16434, A16435, A16437
O2a2b2b1 A16438 , SK1773
O2a2b2b1a SK1775 , SK1776
O2a2b2b1a1 SK1774
O2a2b2b1b A16440 , A16442, A16443, A16444, A16445 O2a3~ M300 O2a4~
M333 J O2b F742 , CTS1754/F4354, CTS5152, CTS6677, F1451,
F1986, F2240, F4161 , F4234 O2b1 F1150 ,
F1578, F1886, F4081, F4191, F4214, F4236, PF4440 O2b1a F837 ,
F1770, F2244, F2247, F3338 O2b1a1 F1025 O2b2 F1055 , F1238, F2051, F2944, F4098, F4178, F4179, F4194, F4238, F4245, F4279 O2b2a F3021 , F4050, F4061, F4323
NOTES:
Identical SNPs that were discovered separately are listed
in alphabetical order, not necessarily in the order of discovery, and
separated by "/". Examples: M307.2/P203.2, M176/SRY465, DYS257, P27.2,
P201/021354
P164 is now defined to be an A to G mutation (rs17316007,
Y-position 12511232) downstream of P201 and upstream of M134. P386 is
now defined to be a T to C mutation downstream of M7. These snps were
earlier confused. The tree is reorganized as of 24 June 2011.
The L1 insertion (also known as LINE1) which appeared in
previous versions of the tree is associated with subclades within
sub-group O3 but has been observed to be absent due to recurrent
deletions so is not included in the current version of the tree.
MSY2.2 was removed from tree because not providing reliable
results.
47z was removed from tree not meeting quality guidelines
and replaced by equivalent CTS713.
M312 was found by Kwon et al. (2015) not to provide results
previously reported.
M162_1, M162_2 removed from tree because not meeting
quality guidelines. M121 mutation information as appearing in Karafet (2008) does not seem correct and was replaced 7 Mar 2017.
Mutations previously on tree but found so far
only in one man or in closely related men : N8, N9, N10, N11
were discovered downstream of M175. Listed 6 Apr 2006.M297 is
downstream from M95 and parallel to M88. Listed 7 August 2012.N4 is
downstream from M113. Listed 7 August 2012.P386 is
downstream from M7. Listed 7 August 2012.P103 is
downstream from IMS-JST002611. Listed 18 July 2013.L681 has been
found within L682. Listed 4 January 2016.L387.1 has
been found within P201, Listed 4 January 2016.
Y-DNA haplogroup O probably originated in
East Asia and later migrated into the South Pacific.The lineage
expanded into Taiwan (high frequency in the aboriginal Taiwanese),
Indonesia,Melanesia, Micronesia, and Polynesia.
O-M122 is the predominant sub-group in China. The O-F265 and
O-PK4 lineages are found in Southeast Asian populations of Malaysia,
Vietnam, Indonesia and southern China. The O-P49 sub-group is of
highfrequency in the Japanese and Korean populations.
References: Capredon et al,Tracing
Arab-Islamic Inheritance in Madagascar: Study of the Y-chromosome and
Mitochondrial DNA in the Antemoro. PLoS ONE 8(11): e80932.
doi:10.1371/journal.pone.0080932, 2013.Chen et al,Y
Chromosome Genotyping and Genetic Structure of Zhuang Populations.
(abstract)Acta Genetica Sinica, 33(12), 1060-72, 2006.Cinnioglu et al,Excavating
Y-chromosome Haplotype Strata in Anatolia. (pdf) Human
Genetics. 114:127-148, 2004.Cox M P & Lahr M M,Y-Chromosome
Diversity Is Inversely Associated with Language Affiliation in
PairedAustronesian- and Papuan-Speaking Communities from Solomon
Islands. (pdf)American Journal of Human Biology, 18:35-50,
2006. Cruciani et al,A Back
Migration from Asia to Sub-Saharan Africa Is Supportedby
High-Resolution Analysis of Human Y-Chromosome Haplotypes. American
Journal of Human Genetics, 70:1197-1214, 2002.Dein et al,The
Y Chromosome Landscape of the Philippines: Extensive Heterogeneity and
Varying Genetic Affinities ofNegrito and Non-Negrito Groups.
(abstract)European Journal of Human Genetics, 19:224-30, 2011.Deng et al,
Evolutionand Migration History of the Chinese Population Inferred from
the Chinese Y-chromosome Evidence. (pdf) Journal of Human
Genetics, 49:339-348, 2004.Dulik et al, Y-Chromosome
Variation in Altaian Kazakhs Reveals a Common Paternal Gene Pool for
Kazakhs and theInfluence of Mongolian Expansions. (pdf)PLoS
ONE, Vol. 6, Issue 3, e17548, 2011.Hallast et al, The
Y-chromosome Tree Bursts into Leaf: 13,000 High-confidence SNPs
Covering the Majority of Known Clades , Molecular Biology and
Evolution, doi: 10.1093/molbev/msu327, 2014.Hammer et al,Dual
Origins of the Japanese: Common Ground for Hunter-gatherer and Farmer Y
Chromosomes. (abstract) Journal of Human Genetics, 51:47-58,
2006.Jin et al,Genetic
Diversity of Two Haploid Markers in the Udegey Population from
Southeastern Siberia. (abstract)American Journal of Physical
Anthropology, 142:303-313, 2010.Jin et al,The
Peopling of Korea Revealed by Analyses of Mitochondrial DNA and
Y-Chromosomal Markers. PLoS One, vol 4(1):ed4210, 2009.Karafet et al,New
Binary Polymorphisms Reshape and Increase Resolution of the Human
Y-Chromosomal HaplogroupTree. Abstract. Genome Research,
published online April 2, 2008.Supplementary
Material. Karafet et al, Paternal
Population History of East Asia: Sources, Patterns,and
Microevolutionary Processes. (pdf) American Journal of Human
Genetics,69:615-628, 2001.Karmin et al, A
Recent Bottleneck of Y chromosome Diversity Coincides with a Global
Change in Culture. Genome Research, doi:
10.1101/gr.186684.114, published online March 13, 2015.Kayser et al, The
Impact of the Austronesian Expansion: Evidence from mtDNa and Y
Chromosome Diversityin the Admiralty Islands of Melanesia. Molecular
Biology Evolution, 25(7):1362-1374, 2008.Kayser et al, Independent
Histories of Human Y Chromosomes from Melanesia and Australia. American
Journal of Human Genetics, 68:173-190, 2001.Kayser et al, Melanesian
and Asian Origins of Polynesians: mtDNA and Y-Chromosome Gradients
acrossthe Pacific. MBE Advance Access published August 21,
2006.Kayser et al. Reduced
Y-Chromosome, but Not Mitochondrial DNA, Diversity in Human Populations
from West NewGuinea. American Journal of Human Genetics,
72:281-302, 2003. Kwon et al. Confirmation
of Y Haplogroup Tree Topologies with Newly Suggested Y-SNPs for the C2,
O2b and O3a Subhaplogroups. (abstract) Forensic Science
International Genetics 19:42-6. doi: 10.1016/j.fsigen.2015.06.003, 19
November 2015. Li et al,Paternal
Genetic Affinity between Western Austronesians and Daic Populations BMC
Evolutionary Biology, Vol. 15(8), p. 146, 2008.Li et al,Substitution
of Hainan Indigenous Genetic Lineage in the Utsat People, Exiles of the
Champa Kingdom Journal of Systematics and Evolution, 51(3),
287-94.Lippold et al,Human
Paternal and Maternal Demographic Histories: Insights from
High-Resolution Y Chromosome and mtDNA Sequences.
Investigative Genetics, 5(13):unpaginated, September 24, 2014.Magoon et al, Generation
of High-resolution a priori Y-chromosome Phylogenies Using
"Next-generation" Sequencing Data bioRxiv, doi:
10.1101/000802, December 13, 2013.Mohyuddin et al,Detection
of Novel Y SNPs Provides Further Insights into Y Chromosomal Variation
in Pakistan. Journal of Human Genetics, 2006.Mona et al,Patterns
of Y-chromosome Diversity Intersect with the Trans-New Guinea
Hypothesis. Mol Biol Evol. 2007 Sep 10; [Epub ahead of print]Naitoh S, et al,Assignment
of Y-chromosomal SNPs Found in Japanese Population to Y-chromosomal
Haplogroup tree .Journal of Human Genetics, 2013 Feb 7. doi:
10.1038/jhg.2012.159, 2013.Ning et al,Refined
Phylogenetic Structure of an Abundant East Asian Y-Chromosomal
Haplogroup O*-M134. (abstract)European Journal of Human
Genetics. (abstract) [EPub ahead of print] doi:10.1038/ejhg.2015.183,
26 August 2015. Nonaka et al,Y
Chromosomal Binary Haplogroups in the Japanese Population and their
Relationship to 16 Y-STR Polymorphisms. (abstract)Annals of
Human Genetics, 71:480-495, 2007. Park et al,Understanding
the Y Chromosome Variation in KoreaRelevance of Combined Haplogroup
and Haplotype Analyses. (abstract)International Journal of
Legal Medicine, 126(4):589-99, 2012. Regueiro et al,Iran:
Tricontinental Nexus for Y-Chromosome Driven Migration.
(abstract)Human Heredity, Vol. 61, No 3, 132-143, 2006.Rootsi S,Human
Y Chromosomal Variations in European Populations .
(dissertation) Council of theInstitute of Molecular and Cell Biology,
University of Tartu, Oct 2004.Rozen et al,Remarkably
Little Variation in Proteins Encoded by the Y Chromosome's Single-Copy
Genes, Implying EffectivePurifying Selection. American
Journal of Human Genetics. 2009 December 11; 85(6): 923-928.Scheinfeldt et al,Unexpected
NRY Chromosome Variation in Northern Island Melanesia. Society
for Molecular Biology, 2006.Sengupta et al,Polarity
and Temporality of High Resolution Y-chromosome Distributions in
IndiaIdentify Both Indigenous and Exogenous Expansions and Reveal Minor
Genetic Influenceof Central Asian Pastoralists. (pdf)American
Journal of Human Genetics, 78:202-221, 2006.Shen et al, Reconstructionof
Patrilineages and Matrilineages of Samaritans and other Israeli
Populations from Y-Chromosomeand Mitochondrial DNA Sequence Variation.
(pdf) Human Mutation, 24:248-260, 2004.Shi et al,Y-Chromosome
Evidence of Southern Origin of the East Asian-SpecificHaplogroup
O3-M122. (abstract)American Journal of Human Genetics.
77:408-419, 2005.Shi Yan et al,An
Updated Tree of Y-chromosome Haplogroup O and Revised Phylogenetic
Positions of Mutations P164 and PK4. European Journal of Human
Genetics advance online publication 20 April 2011; doi:
10.1038/ejhg.2011.64Su et al,Y-chromosome
Evidence for a Northward Migration of Modern Humans into Eastern
Asiaduring the Last Ice Age. (pdf) American Journal of Human
Genetics, 65:1718-1724, 1999.Tan S, et al,Y-chromosome
Polymorphisms Define the Origin of the Mang, an Isolated Population in
China. Annals of Human Biology, Sep;34(5):573-581, 2007Thangaraj et al,Genetic
Affinities of the Andaman Islanders, a Vanishing Human Population.
(pdf)Current Biology, 13:86-93, 2003.Trejaut et al,Taiwan
Y-chromosomal DNA Variation and its Relationship with Island Southeast
Asia. BMC Genetics, 15:77, 2014.Xue Y, et al,Male
Demography in East Asia: A North-South Contrast in Human Population
Expansion Times. (abstract)Genetics, 172:2431-2439, 2006Xue et al,A
Spatial Analysis of Genetic Structure of Human Populations in China
Reveals Distinct Difference betweenMaternal and Paternal Lineages. European
Journal of Human Genetics, 16:705-17, 2008.Wang et al,Late
Neolithic Expansion of Ancient Chinese Revealed by Y chromosome
Haplogroup O3a1c-002611. (abstract)Journal of Systematics and
Evolution, 51:3, 24 Mar 2013.Wang et al,Inferring
Human History in East Asia from Y Chromosomes. Investigative
Genetics, doi: 10.1186/2041-2223-4-11, 2014.Wang et al,Genetic
Structure of Qiangic Populations Residing in the Western Sichuan
Corridor. PLoS One, 9:8:e103772. doi:
10.1371/journal.pone.0103772, 2014.Wei et al, Phylogeography
of Y-chromosome Haplogroup O3a2b2-N6 Reveals Patrilineal Traces of
Austronesian Populations on the Eastern Coastal Regions of Asia . PLoS One, doi:
10.1371/journal.pone.0175080, 2017.
Y
chromosomes of 40% Chinese Descend from Three Neolithic
Super-Grandfathers. PLoS One, 9:8:e103772. doi:
10.1371/journal.pone.0103772, 2014.Yao et al, Improved Phylogenetic Resolution for Y-Chromosome Haplogroup O2a1c-002611 Scientific Reports, 7:1146, doi:10.1038/s41598-017-01340-z, 2017.
Taiwanese
Aborigines: Genetic Heterogeneity and Paternal Contribution to Oceania.
(abstract)Gene, 542(2): 240-47, 2014.Zhang et al,An
Updated Phylogeny of the Human Y-Chromosome Lineage O2a-M95 with Novel
SNPs. PLoS One, 9(6):e101020, 2014.Zhao et al,Presence
of Three Different Paternal Lineages among North Indians: A Study of
560 Y Chromosomes. (abstract)Annals of Human Biology,
36(1):46-59, 2009.Zhong et al,Extended
Y-chromosome Investigation Suggests Post-Glacial Migrations of Modern
Humans into East Asia via the Northern Route. Molecular
Biology, 28(1):717-727, 2011.
Additional Resources: ISOGG
Wiki - What you need to know about Genetic Genealogy.O
Y-Haplogroup Project , Denis Grigoriev,
Sastrosudharmo, Owen Lu.
Corrections/Additions made since 1 January 2017:
Added F13, F30,
F62, F68, F251, F253, F379, F514, F530, F60, F325, F362, F526,
F561, FGC12472/Y3271/Z25059 , CTS2107,
CTS2128, CTS2916, CTS3518, CTS4150, CTS8491, CTS10670, SK1691, SK1692,
SK1695, SK1696, PH203, FGC1803.2/Y2044.2, PH278, PH975, PH1787, PH2385,
PH3699, PH3838, PH3849, PH4404, PH4535, PH4997, PH5031, PH5323 to tree;
updated additional resources on 2 February 2017.
Added A12446, A12447, A12448, PH40, PH437 on 7 February
2017.
Added FGC54466, FGC54513, FGC54523, FGC54527, Y15977,
Y26385, Added CTS10944, CTS12052, Z25245, Y27939, Y27942, Y27943, Y27951, ZS2079 to tree on 15 February 2017. Added Y26403,
Y26409, Y26412, Y26413, Y26415, Y26416, Y26421, Y26422, Y26424, Y26428,
Y26430, Y26433, Y26434, Y26437, Y26440, Y26446, Y26448, Y26457, Y26460,
Y27924, Y27927, Y27932, Y27933, Y27934 to tree on 16 February 2017. Renamed F1316 to F1316C to reflect revised name and corresponding mutation information on 19 February 2017.. Added
Z25061, CTS362, CTS2801, CTS6777, CTS9127, F64, F92, F691, Z25075,
Z25076, Z25077, Z25079, Z25081, Y20951, Z43887, F1418, F1495, Z25097, Z25098, Z43869, Z43870, Z43871, Z43872, Z43873, Z43874, Z43875, Z43876, Z43877, Z43878, Z43879, Z43887, Z43880. Z43881, Z43882, Z43883, Z43884, Z43885, Z43886 to tree on 20 February 2017.Added F117, FGC12471/Y3268. CTS5820 to tree on 22 February 2017. Added CTS498,
CTS957, CTS1028, CTS2398, CTS4941, CTS5926, CTS7341, CTS8263, CTS9516,
CTS10139, CTS10185, CTS11076, CTS11949 to tree on 24 February 2017. Added CTS3709, CTS8168, CTS8587, CTS10730, F852, F1054, F4348/Z24834, SK1669/Z24838, Z24836, Z24840, Z24844, Z43961,
Z43962, Z43963, Z43964, Z43965, Z43966, Z43967, Z43968,
Z43969, Z43970, Z43971, Z43972, Z43973, Z43974, Z43975, Z43976,
Z43977, Z43978, Z43979, Z43980 to tree; revised mutation information
for M121 to show correct information on 8 March 2017.Added Y29828, Y30477, Y30485, Y30487, Y30489, Y30493, Y30498, ZS11791, F735, L932.2, Y29836, Y30471, Y30484, Y30499, FGC34973, Y29829, Y29830, Y29831, Y29832, Y29833, Y29834, Y29835,
Y30465, Y30466, Y30467, Y30468, Y30469, Y30470, Y30472, Y30473, Y30474,
Y30475, Y30476, Y30478, Y30479, Y30480, Y30481, Y30482, Y30483, Y30486,
Y30488, Y30490, Y30491, Y30492, Y30494, Y30495, Y30496, Y30497, Y30500 to tree on 11 March 2017. Added YP4864,
YP4865, Z44052, Z44053, Z44054, Z44055, Z44056, Z44057, Z44058, Z44059,
Z44060, Z44061, Z44062, Z44063, Z44064, Z44065, Z44066, Z44067 , Z44068, Z44069, Z44071, Z44072,
Z44073, Z44074, Z44075, Z44076, Z44077, Z44078, Z44079, Z44080, Z44081,
Z44082, Z44083, Z44084, Z44085, Z44086, Z44087, Z44088, Z44089, Z44090, Z44091, Z44092, Z44093, Z44094, Y29861, Y30150 to tree; added F5524/SK1746, Z44070 , F5525/SK1748 to tree investigation on 16 March 2017. Added F1442, F2779, F1123, F2265, F2425, F2453, F3048, F3115 to tree; added F1369 to tree investigation on 17 March 2017. Added CTS1754/F4354, CTS5152, CTS6677, F1451, F2240, F4234, F1578, F1886, F4081, F4191, F4214, PF4440, F1025, F1055, F1238, F2051, F2944, F4098, F4178, F4179, F4194, F4238, F4245, F4279 , F3021, F4050, F4061, F4323 to tree on 21 March 2017.Added SK1780, SK1781, B451/AM01847, AM01781/S18967/SK1786, AM01761/SK1787, AM01798/SK1788 to tree on 11 May 2017. Added Wei et al., Yao et al. to bibliography on 12 May 2017. Added F14400/Z26072, F14403/Z26073, F14487/Z26074, F14497/Z26075,
F14503/Z26076, F14559/Z26077, F14690/Z26080, F14695/Z26081,
F14714/Z26083, FGC61160, FGC61167, Y31256, Z26078, CTS2205, FGC61159, FGC61164, FGC61165, FGC61215, Y31257 , A16615, A16616, A16617,
A16618, A16621, A16622, A16623, A16624, A16625, A16626, A16628, CTS496,
CTS724, CTS1707, CTS2147, CTS2390, CTS8451, CTS9362, Added
A16631, A16632, S24099.2, A16636, A16637, A16638, A16639, A16640,
A16641, A16642, A16643, A16644, A16645 to tree on 15 June 2017. Added A16611, A16612, A16613, CTS6105, PH2506, PH2797, PH5023 to tree on 17 June 2017. Added CTS12480, F4215, F20220, F20394, F24749, Z44734, Z44735, Z44736, Z44737, Z44738, Z44739, Added Z44740, Z44741, Z44742, Z44743, Z44744, Z44745, Z44746, Z44747, Z44748,
Z44749, Z44750, Z44751, Z44752, Z44753, Z44754, Z44755, Z44756, Z44757,
Z44758, Z44759, Z44760, Z44761, Z44762, Z44763, Z44764 to tree on 20 June 2017. Added BY15188, F14875, F15739, F15692, F15973, F16170, F17339, F18455,
F18521, F18757, F19055, F20227, F20235, F20251, F20767, F21091, F21275,
F22909, F23173, F23473, F23524, F23679, F24517, F24605, F25036, F25041,
F25079, F25381, F25411, F25440, F25789, F25924, F26193, F26484, F26752,
Z44765, Z44766, Z44767, Z44768, Z44769, Z44770, Z44771, Z44772, Z44773,
Z44774, Z44775, Z44776, Z44777, Z44778, Z44779, Z44780, Z44781, Z44782,
Z44783, Z44784, Z44785, Z44786, Z44787 to tree on 21 June 2017. Added F16411, F17835, F19270, F19578, F22244, F22838, F23276, F23329, F24971, Z44792 to tree on 22 June 2017. Added Z25701, Z25708, Z25713, Z25716, Z25720, Z25721, Z25728, Z25730, Z25737, Z44816, AM01755, AM01768, AM01780, AM01790/F2486, AM01791, AM01792, AM01799/F2654, AM01801, AM01804, AM01842, AM01857, F16175, Z44817 to tree on 27 June 2017. Added F1010,
F1375, F1412, F1884, F1889, F2132, F2511, F2558. F2724, F2929, F3066,
F3095, F3230, F3473, F3820, SK1782, SK1784, F3612, F3819,
SK1783, AM01750, AM01759, AM01760, AM01762, AM01763, AM01771,
AM01775, AM01778,
AM01785, AM01786, AM01787, AM01794, AM01802, AM01805, AM01806, AM01812,
AM01813, AM01815, AM01818, AM01821, AM01824, AM01827, AM01835, AM01838,
AM01839, AM01844, AM01855, AM01859, AM01860, F15980, F16102, F16868,
F17803, A17418, A17419, A17420, A17422, A17424, A17425, A17426, A17427,
A17428, A17429, A17430, A17431, A17432, A17433, A17434, A17436 to tree
on 28 June 2017.Added
A17437, A17438, A17439, A17441, A17442, A17443, A17444, A17445, A17446,
A17447, A17448, A17449, A17450, A17451, YP6194, AM01756, AM01757,
AM01779, AM01803, AM01810, AM01826, AM01834, AM01861, F16663, F16771,
F16783, B450, AM00474/SK1791, AM00475/SK1792, AM00476,
F16769, AM00472/B452, AM00463, AM00464, AM00465, AM00466, AM00467,
AM00468,
AM00469, AM00470, AM00471, F16088, F16616, F16997, F17764,
F26682, F18942, F21615, F25531, A16427, A16428, A16429,
A16430, A16431, A16432, A16433, A16434, A16435,
A16437, A16438, SK1773, SK1775, SK1776, SK1774, A16440,
A16442, A16443, A16444, A16445 to tree on 29 June 2017. Added
F20963, Z44926, Z44927, M1513, M1557, M1593, M1602, M1604, M1608,
M1617, M1629, M1637, M1683, M1688, M1719, Z44923, Z44924, Z44925,
F16367, F24592, F17158, Z44918, Z44919, Z44921, Z44922 to tree on
7 July 2017. Added
F4212, F4251, FGC61079, FGC61080, PF3452, Z23746, Z23781, Z23787,
Z23799, F809, Y31259, Y32109, Z23745, Z23766, Z23777, Z23784,
Z23786, Z23791, Z23797, Z23801, Z23803, Z23804, F2517 to tree on 14
July 2017. Added FGC19670/Y14253/Z23825, FGC19672/Y14260/Z23840,
FGC19673/SK1620/Y14000/Z23821, FGC19674/Y14018/Z23835,
FGC19675/Y13998/Z23819, FGC19676/Y14255/Z23831, FGC19677/Y14259/Z23834,
FGC19678/Y14257, FGC19679/Y14008/Z23828, FGC19681/Y14020/Z23836,
FGC19682/Y14261/Z23841, FGC19683/Y14011/Z23829, FGC19684/Y14014/Z23832,
FGC19686/Y14022/Z23839, FGC19687/Y13999/Z23820, FGC19688/Y14007/Z23827,
FGC19689/SK1619/Y14001/Z23822, FGC19694/SK1618/Y14256, FGC19705/Y14010,
FGC19710/Y14021/Z23837, Z23824, Z23833, Z23838, Z23843, FGC19671/Z23845, FGC19690/Y14251/Z23851, FGC19693/Y14013/Z23857,
FGC19698/Y13996/Z23852, Z23849, FGC19700/Y14017/Z23861, FGC19701/Y14002/Z23853,
FGC19703/Y14262/Z23866, FGC19711/Y14025/Z23865, FGC19712/Y14016/Z23860, FGC19715/Y13997, FGC19724, FGC61038, FGC61039, FGC61054, Y31276, Y31927, Y31928
