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Coefficient of relationship

From ISOGG Wiki

The coefficient of relationship is a measure of the degree of consanguinity (or biological relationship) between two individuals. The term coefficient of relationship was defined by Sewall Wright in 1922, and was derived from his definition of the coefficient of inbreeding of 1921. The measure is most commonly used in genetics and genetic genealogy.

In general, the higher the level of inbreeding the closer the coefficient of relationship approaches a value of 1, expressed as a percentage, and approaches a value of 0 for individuals with arbitrarily remote common ancestors.[1]

In human relationships, the value of the coefficient of relationship is usually calculated based on the knowledge of a full family tree extending to a comparatively small number of generations, perhaps of the order of three or four. As explained above, the value for the coefficient of relationship so calculated is thus a lower bound, with an actual value that may be up to a few per cent higher. The value is accurate to within 1% if the full family tree of both individuals is known to a depth of seven generations.[2]

Relationship Coefficient of
relationship (r)
Identical twins; clones 100%[3] (1)
Parent-offspring 50% (2−1)
Full siblings 50% (2−2+2−2)
3/4 siblings or sibling-cousins 37.5% (2−2+2−3)
Grandparent-grandchild 25% (2−2)
Half siblings 25% (2−2)
Aunt/uncle-nephew/niece 25% (2⋅2−3)
Double first cousins 25% (4⋅2−4)
Great grandparent-great grandchild 12.5% (2−3)
First cousins 12.5% (2⋅2−4)
Quadruple second cousins 12.5% (8⋅2−6)
Triple second cousins 9.38% (6⋅2−6)
Half-first cousins 6.25% (2−4)
First cousins once removed 6.25% (2⋅2−5)
Double second cousins 6.25% (4⋅2−6)
Second cousins 3.13% (2⋅2−6)
Third cousins 0.78% (2⋅2−8)
Fourth cousins 0.20% (2⋅2−10)[4]

Further reading

Licence

This article is licensed under the GNU Free Documentation Licence. It uses material from the Wikipedia article Coefficient of relationship.

References

  1. Strictly speaking, r=1 for clones and identical twins, but since the definition of r is usually intended to estimate the suitability of two individuals for breeding, they are typically taken to be of opposite sex.
  2. A full family tree of seven generations (128 paths to ancestors of the 7th degree) is unreasonable even for members of high nobility. For example, the family tree of Queen Elizabeth II is fully known for a depth of six generations, but becomes difficult to trace in the seventh generation.
  3. By replacement in the definition of the notion of "generation" by meiosis". Since identical twins are not separated by meiosis, there are no "generations" between them, hence n=0 and r=1. See genetic-genealogy.co.uk.
  4. This degree of relationship is usually indistinguishable from the relationship to a random individual within the same population (tribe, country, ethnic group).